Achlya

Achlya is a genus of oomycete (water mold), in the family Saprolegniaceae. The genus includes several plant pathogens including Achlya conspicua and Achlya klebsiana. Unlike many other microorganisms, cell expansion is governed by changes in cell wall strength rather than changes in osmotic pressure. The genome of Achlya hypogyna has been sequenced and can be accessed on public online databases, for example on the NCBI website (National Center for Biotechnology Information).

The genus was first circumscribed in Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. vol.11 on page 514 in 1823 by Christian Gottfried Daniel Nees von Esenbeck.

The type species is Achlya prolifera.

The genus name of Achlya is named after Achlys, who in the Hesiodic Shield of Heracles, is one of the figures depicted on Heracles' shield, perhaps representing the personification of sorrow.

Several Achyla spp. are pathogens of rice seedlings.

Species
As accepted by Species Fungorum;


 * Achlya abortispora
 * Achlya abortiva
 * Achlya achlyoides
 * Achlya ambisexualis Raper (1939)
 * Achlya ambispora Steciow (2001)
 * Achlya americana
 * Achlya androcomposita
 * Achlya anomala
 * Achlya aquatica Dayal & J. Thakur (1969)
 * Achlya bisexualis Coker & Couch (1927)
 * Achlya bispora
 * Achlya bonariensis
 * Achlya caroliniana
 * Achlya colorata Pringsh. (1882)
 * Achlya conspicua Coker (1923)
 * Achlya crenulata
 * Achlya debaryana
 * Achlya diffusa
 * Achlya dubia
 * Achlya echinulata
 * Achlya flagellata
 * Achlya flexuosa
 * Achlya formosana
 * Achlya fuegiana
 * Achlya glomerata
 * Achlya haehneliana
 * Achlya heterosexualis
 * Achlya imperfecta Coker (1923)
 * Achlya inflata
 * Achlya intricata
 * Achlya kamatii
 * Achlya kashyapia
 * Achlya klebsiana Pieters (1915)
 * Achlya lobata
 * Achlya michiganensis
 * Achlya orion Coker & Couch (1920)
 * Achlya oryzae
 * Achlya oviparvula
 * Achlya pacifica
 * Achlya papillosa
 * Achlya primoachlya
 * Achlya prolifera
 * Achlya proliferoides
 * Achlya pseudoachlyoides
 * Achlya pseudoradiosa
 * Achlya racemosa Hildebr. (1867)
 * Achlya radiosa
 * Achlya regularis
 * Achlya robusta
 * Achlya rodrigueziana
 * Achlya sparrowii
 * Achlya spiracaulis
 * Achlya spiralis
 * Achlya subterranea
 * Achlya truncatiformis M.W. Dick & Mark A. Spencer (2002)
 * Achlya tuberculata
 * Achlya turfosa

Former species; (all are Saprolegniaceae family)


 * A. abortiva f. normalis = Achlya abortiva
 * A. acadiensis = Newbya androgyna,
 * A. ambisexualis var. abjointa = Achlya ambisexualis
 * A. ambisexualis var. gracilis = Achlya ambisexualis
 * A. americana var. cambrica = Achlya americana
 * A. americana var. megasperma = Achlya americana
 * A. americana var. megasperma = Achlya americana
 * A. androgyna = Newbya androgyna
 * A. apiculata = Newbya apiculata
 * A. apiculata var. forbesiana = Newbya apiculata
 * A. apiculata var. prolifica = Newbya apiculata
 * A. benekei = Protoachlya benekei
 * A. bisexualis var. ambisexualis = Achlya ambisexualis
 * A. brasiliensis = Newbya brasiliensis
 * A. braunii = Newbya androgyna
 * A. cambrica = Achlya americana
 * A. curvicollis = Newbya curvicollis
 * A. debaryana var. americana = Achlya americana,
 * A. debaryana var. intermedia = Achlya debaryana
 * A. dioica = Saprolegnia dioica, Saprolegniaceae
 * A. dubia var. pigmenta = Achlya dubia
 * A. flagellata var. yezoensis = Achlya flagellata
 * A. klebsiana var. indica = Achlya klebsiana
 * A. lignicola = Achlya racemosa
 * A. megasperma = Newbya megasperma
 * A. mucronata = Protoachlya mucronata
 * A. oblongata = Newbya oblongata
 * A. oblongata var. gigantica = Newbya oblongata
 * A. oblongata var. globosa = Newbya oblongata
 * A. oligocantha = Newbya oligocantha
 * A. oligocantha var. brevispina = Newbya oligocantha
 * A. ornata = Newbya pascuicola
 * A. paradoxa = Isoachlya paradoxa
 * A. polyandra sensu = Achlya debaryana
 * A. polyandra = Newbya polyandra
 * A. racemosa f. maxima = Achlya racemosa
 * A. racemosa f. polyspora = Achlya racemosa
 * A. racemosa var. lignicola = Achlya racemosa
 * A. racemosa var. maxima = Achlya racemosa
 * A. racemosa var. spinosa = Newbya spinosa
 * A. racemosa var. stelligera = Achlya racemosa
 * A. recurva = Newbya recurva
 * A. spinosa = Newbya spinosa
 * A. stellata = Newbya stellata
 * A. stellata var. multispora = Protoachlya mucronata
 * A. treleaseana = Newbya androgyna