Alpheus (crustacean)

Alpheus is a genus of snapping shrimp of the family Alpheidae. This genus contains in excess of 330 species, making this the most species-rich genus of shrimp. Like other snapping shrimp, the claws of Alpheus are asymmetrical, with one of the claws enlarged for making a popping noise. Some species in the genus enter into symbiotic relationships with gobiid fishes, and others associate with sea anemones.

Species
Valid species as of October 2023:
 * Alpheus abumusa Dehghani, Sari & Naderloo, 2019
 * Alpheus acutocarinatus De Man, 1909
 * Alpheus acutofemoratus Dana, 1852
 * Alpheus adamastor Coutière, 1908
 * Alpheus aequus Kim & Abele, 1988
 * Alpheus africanus Balss, 1916
 * Alpheus agilis Anker, Hurt & Knowlton, 2009
 * Alpheus agrogon Ramos, 1997
 * Alpheus alaincrosnieri Anker, 2020
 * Alpheus albatrossae (AH Banner, 1953)
 * Alpheus alcyone De Man, 1902
 * Alpheus alpheopsides Coutière, 1905
 * Alpheus amarillo Anker, 2012
 * Alpheus amblyonyx Chace, 1972
 * Alpheus amirantei Coutière, 1908
 * Alpheus anchistus De Man, 1920
 * Alpheus angulosus McClure, 2002
 * Alpheus angustilineatus Nomura & Anker, 2005
 * Alpheus ankeri Dehghani, Sari & Naderloo, 2019
 * Alpheus antepaenultimus Kim & Abele, 1988
 * Alpheus architectus De Man, 1897
 * Alpheus arenensis (Chace, 1937)
 * Alpheus arenicolus Banner & Banner, 1983
 * Alpheus arethusa De Man, 1909
 * Alpheus armatus Rathbun, 1901
 * Alpheus armillatus H. Milne Edwards, 1837
 * Alpheus arnoa Banner, 1957
 * Alpheus astrinx Banner & Banner, 1982
 * Alpheus australiensis Banner & Banner, 1982
 * Alpheus australosulcatus Banner & Banner, 1982
 * Alpheus baccheti Anker, 2010
 * Alpheus bahamensis Rankin, 1898
 * Alpheus balaenodigitus Banner & Banner, 1982
 * Alpheus bannerorum Bruce, 1987
 * Alpheus barbadensis (Schmitt, 1924)
 * Alpheus barbatus Coutière, 1897
 * Alpheus batesi Banner & Banner, 1964
 * Alpheus bellimanus Lockington, 1877
 * Alpheus bellulus Miya & Miyake, 1969
 * Alpheus bicostatus De Man, 1908
 * Alpheus bidens (Olivier, 1811)
 * Alpheus bisincisus De Haan, 1849
 * Alpheus blachei Crosnier & Forest, 1965
 * Alpheus bouvieri A. Milne-Edwards, 1878
 * Alpheus brachymerus (AH Banner, 1953)
 * Alpheus bradypus Coutière, 1905
 * Alpheus brasileiro Anker, 2012
 * Alpheus brevicristatus De Haan, 1844
 * Alpheus brevipes Stimpson, 1860
 * Alpheus brevirostris (Olivier, 1811)
 * Alpheus brucei Banner & Banner, 1982
 * Alpheus bucephaloides Nobili, 1905
 * Alpheus bucephalus Coutière, 1905
 * Alpheus buchanorum Banner & Banner, 1983
 * Alpheus buckupi Almeida, Terossi, Araújo-Silva & Mantelatto, 2013
 * Alpheus bunburius Banner & Banner, 1982
 * Alpheus burukovskyi Anker & Pachelle, 2015
 * Alpheus californiensis Holmes, 1900
 * Alpheus canaliculatus Banner & Banner, 1968
 * Alpheus candei Guérin-Méneville, 1855
 * Alpheus carlae Anker, 2012
 * Alpheus cedrici Anker & De Grave, 2012
 * Alpheus chacei Carvacho, 1979
 * Alpheus chamorro Banner, 1956
 * Alpheus chilensis Coutière in Lenz, 1902
 * Alpheus chiragricus H. Milne Edwards, 1837
 * Alpheus christofferseni Anker, Hurt & Knowlton, 2007
 * Alpheus clamator Lockington, 1877
 * Alpheus clypeatus Coutière, 1905
 * Alpheus coetivensis Coutière, 1908
 * Alpheus collumianus Stimpson, 1860
 * Alpheus colombiensis Wicksten, 1988
 * Alpheus compressus Banner & Banner, 1981
 * Alpheus confusus Carvacho, 1989
 * Alpheus coutierei De Man, 1909
 * Alpheus crinitus Dana, 1852
 * Alpheus cristatus Coutière, 1897
 * Alpheus cristulifrons Rathbun, 1900
 * Alpheus crockeri (Armstrong, 1941)
 * Alpheus cylindricus Kingsley, 1878
 * Alpheus cythereus Banner & Banner, 1966
 * Alpheus dasycheles Coutière, 1908
 * Alpheus davaoensis Chace, 1988
 * Alpheus dentipes Guérin, 1832
 * Alpheus deuteropus Hilgendorf, 1879
 * Alpheus diadema Dana, 1852
 * Alpheus digitalis De Haan, 1844
 * Alpheus dingabadi Anker, 2023
 * Alpheus dispar Randall, 1840
 * Alpheus distinctus Kim & Abele, 1988
 * Alpheus djeddensis Coutière, 1897
 * Alpheus djiboutensis De Man, 1909
 * Alpheus dolerus Banner, 1956
 * Alpheus echiurophilus Anker, Komai & Marin, 2015
 * Alpheus edamensis De Man, 1888
 * Alpheus edwardsii (Audouin, 1826)
 * Alpheus ehlersii De Man, 1909
 * Alpheus estuariensis Christoffersen, 1984
 * Alpheus euchirus Dana, 1852
 * Alpheus eulimene De Man, 1909
 * Alpheus euphrosyne De Man, 1897
 * Alpheus eurydactylus De Man, 1920
 * Alpheus exilis Kim & Abele, 1988
 * Alpheus explorator Boone, 1935
 * Alpheus facetus De Man, 1908
 * Alpheus fagei Crosnier & Forest, 1965
 * Alpheus fasciatus Lockington, 1878
 * Alpheus fasqueli Anker, 2001
 * Alpheus felgenhaueri Kim & Abele, 1988
 * Alpheus fenneri Bruce, 1994
 * Alpheus firmus Kim & Abele, 1988
 * Alpheus floridanus Kingsley, 1878
 * Alpheus foresti Banner & Banner, 1981
 * Alpheus formosus Gibbes, 1850
 * Alpheus frontalis H. Milne Edwards, 1837
 * Alpheus fujitai Nomura & Anker, 2005
 * Alpheus funafutensis Borradaile, 1899
 * Alpheus fushima Nomura, 2009
 * Alpheus galapagensis Sivertsen, 1933
 * Alpheus gallicus Scioli & Anker, 2020
 * Alpheus georgei Banner & Banner, 1982
 * Alpheus glaber (Olivi, 1792)
 * Alpheus gracilipes Stimpson, 1860
 * Alpheus gracilis Heller, 1861
 * Alpheus grahami Abele, 1975
 * Alpheus haanii Ortmann, 1890
 * Alpheus hailstonei Coutière, 1905
 * Alpheus halesii Kirk, 1887
 * Alpheus hebes Kim & Abele, 1988
 * Alpheus heeia Banner & Banner, 1975
 * Alpheus hephaestus Bracken-Grissom & Felder, 2014
 * Alpheus heronicus Banner & Banner, 1982
 * Alpheus heterocarpus (Yu, 1935)
 * Alpheus heterochaelis Say, 1818
 * Alpheus heurteli Coutière, 1897
 * Alpheus hippothoe De Man, 1888
 * Alpheus holthuisi Ribeiro, 1964
 * Alpheus homochirus (Yu, 1935)
 * Alpheus hoonsooi Kim & Abele, 1988
 * Alpheus hoplocheles Coutière, 1897
 * Alpheus hortensis Wicksten & McClure, 2003
 * Alpheus hululensis Coutière, 1905
 * Alpheus hutchingsae Banner & Banner, 1982
 * Alpheus hyeyoungae Kim & Abele, 1988
 * Alpheus hyphalus Chace, 1988
 * Alpheus idiocheles Coutière, 1905
 * Alpheus ikedosoma Komai, 2015
 * Alpheus imitatrix De Man, 1909
 * Alpheus immaculatus Knowlton & Keller, 1983
 * Alpheus inca Wicksten & Méndez G., 1981
 * Alpheus inopinatus Holthuis & Gottlieb, 1958
 * Alpheus intrinsecus Spence Bate, 1888
 * Alpheus japonicus Miers, 1879
 * Alpheus javieri Anker, Hurt & Knowlton, 2009
 * Alpheus kagoshimanus Hayashi & Nagata, 2000
 * Alpheus karplusi Anker, 2022
 * Alpheus kuroshimensis Nomura & Anker, 2005
 * Alpheus labis Banner & Banner, 1982
 * Alpheus lacertosus Kim & Abele, 1988
 * Alpheus ladronis Banner, 1956
 * Alpheus lanceloti Coutière, 1905
 * Alpheus lanceostylus Banner, 1959
 * Alpheus lancirostris Rankin, 1900
 * Alpheus latus Kim & Abele, 1988
 * Alpheus lentiginosus Anker & Nizinski, 2011
 * Alpheus lepidus De Man, 1908
 * Alpheus leptocheles Banner & Banner, 1975
 * Alpheus leptochiroides De Man, 1909
 * Alpheus leptochirus Coutière, 1905
 * Alpheus leviusculus Dana, 1852
 * Alpheus lobidens De Haan, 1849
 * Alpheus longecarinatus Hilgendorf, 1879
 * Alpheus longichaelis Carvacho, 1979
 * Alpheus longiforceps Hayashi & Nagata, 2002
 * Alpheus longinquus Kim & Abele, 1988
 * Alpheus longipalma Komai & Ohtomi, 2018
 * Alpheus lottini Guérin, 1829
 * Alpheus luiszapatai Ramos-Tafur, 2018
 * Alpheus lutosus Anker & De Grave, 2009
 * Alpheus macellarius Chace, 1988
 * Alpheus macrocheles (Hailstone, 1835)
 * Alpheus macrodactylus Ortmann, 1890
 * Alpheus macroskeles Alcock & Anderson, 1899
 * Alpheus maindroni Coutière, 1898
 * Alpheus malabaricus (Fabricius, 1775)
 * Alpheus malleator Dana, 1852
 * Alpheus malleodigitus (Spence Bate, 1888)
 * Alpheus mangalis Anker, 2023
 * Alpheus mannarensis Purushothaman, Abhilash, Ajith Kumar & Lal, 2021
 * Alpheus margaritae Salgado-Barragán, Ayón-Parente & Zamora-Tavares, 2017
 * Alpheus martini Kim & Abele, 1988
 * Alpheus mathewsae Anker, 2012
 * Alpheus mazatlanicus Wicksten, 1983
 * Alpheus microrhynchus De Man, 1897
 * Alpheus microscaphis (Banner, 1959)
 * Alpheus microstylus (Spence Bate, 1888)
 * Alpheus miersi Coutière, 1898
 * Alpheus migrans Lewinsohn & Holthuis, 1978
 * Alpheus millsae Anker, Hurt & Knowlton, 2007
 * Alpheus mitis Dana, 1852
 * Alpheus mohammadpouri Dehghani, Sari & Naderloo, 2019
 * Alpheus moretensis Banner & Banner, 1982
 * Alpheus naos Anker, Hurt & Knowlton, 2007
 * Alpheus naranjo Anker, 2018
 * Alpheus nipa Banner & Banner, 1985
 * Alpheus nobili Banner & Banner, 1966
 * Alpheus nomurai Anker, 2023
 * Alpheus nonalter Kensley, 1969
 * Alpheus normanni Kingsley, 1878
 * Alpheus notabilis Stebbing, 1915
 * Alpheus novaezealandiae Miers, 1876
 * Alpheus nuno Anker, 2012
 * Alpheus nuttingi (Schmitt, 1924)
 * Alpheus oahuensis (AH Banner, 1953)
 * Alpheus obesomanus Dana, 1852
 * Alpheus octocellatus Anker & Benzoni, 2023
 * Alpheus ovaliceps Coutière, 1905
 * Alpheus pachychirus Stimpson, 1860
 * Alpheus pacificus Dana, 1852
 * Alpheus packardii Kingsley, 1880
 * Alpheus paludicola Kemp, 1915
 * Alpheus panamensis Kingsley, 1878
 * Alpheus papillosus Banner & Banner, 1982
 * Alpheus paracrinitus Miers, 1881
 * Alpheus paradentipes Coutière, 1905
 * Alpheus paraformosus Anker, Hurt & Knowlton, 2008
 * Alpheus paralcyone Coutière, 1905
 * Alpheus paralpheopsides Coutière, 1905
 * Alpheus parasocialis Banner & Banner, 1982
 * Alpheus pareuchirus Coutière, 1905
 * Alpheus parvimaculatus Nomura & Anker, 2005
 * Alpheus parvirostris Dana, 1852
 * Alpheus parvus De Man, 1909
 * Alpheus peasei (Armstrong, 1940)
 * Alpheus percyi Coutière, 1908
 * Alpheus perezi Coutière, 1908
 * Alpheus perlas Anker & Pachelle, 2019
 * Alpheus perplexus Banner, 1956
 * Alpheus petronioi Almeida, Terrosi & Mantelatto, 2014
 * Alpheus philoctetes De Man, 1909
 * Alpheus platycheirus Boone, 1927
 * Alpheus platydactylus Coutière, 1897
 * Alpheus platyunguiculatus (AH Banner, 1953)
 * Alpheus polystictus Knowlton & Keller, 1985
 * Alpheus polyxo De Man, 1909
 * Alpheus pontederiae de Rochebrune, 1883
 * Alpheus pouang Christoffersen, 1979
 * Alpheus praedator De Man, 1908
 * Alpheus proseuchirus De Man, 1908
 * Alpheus pseudopugnax (AH Banner, 1953)
 * Alpheus puapeba Christoffersen, 1979
 * Alpheus pubescens De Man, 1908
 * Alpheus pugnax Dana, 1852
 * Alpheus punctatus Anker, 2012
 * Alpheus pustulosus Banner & Banner, 1968
 * Alpheus quasirapacida Chace, 1988
 * Alpheus ramosportoae Soledade, Terossi, Scioli, Mantelatto & Almeida, 2019
 * Alpheus randalli Banner & Banner, 1980
 * Alpheus rapacida De Man, 1908
 * Alpheus rapax Fabricius, 1798
 * Alpheus rectus Kim & Abele, 1988
 * Alpheus ribeiroae Anker & Dworschak, 2004
 * Alpheus richardsoni Yaldwyn, 1971
 * Alpheus richpalmeri Anker, 2020
 * Alpheus roblesi Bracken-Grissom & Felder, 2014
 * Alpheus romensky Burukovsky, 1990
 * Alpheus roquensis Knowlton & Keller, 1985
 * Alpheus roseodigitalis Nomura & Anker, 2005
 * Alpheus rostratus Kim & Abele, 1988
 * Alpheus rudolphi Almeida & Anker, 2011
 * Alpheus rugimanus A. Milne-Edwards, 1878
 * Alpheus samoa Banner & Banner, 1966
 * Alpheus samudra De Grave, Krishnan, Kumar K.P. & Christodoulou, 2020
 * Alpheus savuensis De Man, 1908
 * Alpheus saxidomus Holthuis, 1980
 * Alpheus schmitti Chace, 1972
 * Alpheus sciolii Anker, 2022
 * Alpheus scopulus Kim & Abele, 1988
 * Alpheus serenei Tiwari, 1964
 * Alpheus sibogae De Man, 1908
 * Alpheus simus Guérin-Méneville, 1855
 * Alpheus sizou Banner & Banner, 1967
 * Alpheus socialis Heller, 1862
 * Alpheus soelae Banner & Banner, 1986
 * Alpheus songkla Banner & Banner, 1966
 * Alpheus soror Bruce, 1999
 * Alpheus spatulatus Banner & Banner, 1968
 * Alpheus splendidus Coutière, 1897
 * Alpheus spongiarum Coutière, 1897
 * Alpheus stanleyi Coutière, 1908
 * Alpheus stantoni Banner & Banner, 1986
 * Alpheus staphylinus Coutière, 1908
 * Alpheus stephensoni Banner & Smalley, 1969
 * Alpheus strenuus Dana, 1852
 * Alpheus styliceps Coutière, 1905
 * Alpheus sudara Banner & Banner, 1966
 * Alpheus sulcatus Kingsley, 1878
 * Alpheus sulcipalma Purushothaman, Bharathi, Damodhar, Kumar & Lal, 2023
 * Alpheus suluensis Chace, 1988
 * Alpheus supachai Banner & Banner, 1966
 * Alpheus superciliaris Coutière, 1905
 * Alpheus takla Anker, 2023
 * Alpheus talismani Coutière, 1898
 * Alpheus tampensis Anker, 2012
 * Alpheus tasmanicus Banner & Banner, 1982
 * Alpheus tenuicarpus De Man, 1908
 * Alpheus tenuipes De Man, 1910
 * Alpheus tenuis Kim & Abele, 1988
 * Alpheus thomasi Hendrix & Gore, 1973
 * Alpheus thompsoni Anker, 2022
 * Alpheus tirmiziae Kazmi, 1974
 * Alpheus tricolor Anker, 2001
 * Alpheus triphopus Nobili, 1906
 * Alpheus tungii Banner & Banner, 1966
 * Alpheus ulalae Bracken-Grissom & Felder, 2014
 * Alpheus umbo Kim & Abele, 1988
 * Alpheus utriensis Ramos & von Prahl, 1989
 * Alpheus vanderbilti Boone, 1930
 * Alpheus vanuatu Anker, 2020
 * Alpheus verrilli (Schmitt, 1924)
 * Alpheus villosus (Olivier, 1811)
 * Alpheus villus Kim & Abele, 1988
 * Alpheus viridari (Armstrong, 1949)
 * Alpheus viserion Anker, Leray & Pachelle, 2021
 * Alpheus vladivostokiensis (Vinogradov, 1950)
 * Alpheus waltervadi Kensley, 1969
 * Alpheus websteri Kingsley, 1880
 * Alpheus wickstenae Christoffersen & Ramos, 1987
 * Alpheus williamsi Bruce, 1994
 * Alpheus wonkimi Anker & Pachelle, 2013
 * Alpheus xanthocarpus Anker, Hurt & Knowlton, 2008
 * Alpheus xishaensis Liu & Lan, 1980
 * Alpheus zarenkovi Anker & Pachelle, 2015
 * Alpheus zimmermani Anker, 2007
 * Alpheus zulfaquiri Kazmi, 1982