Andricus

Andricus is a genus of oak gall wasps in the family Cynipidae.

Life cycle
As in all Hymenoptera, sex-determination in species of the genus Andricus is governed by haplodiploidy: males develop from unfertilized eggs and are haploid, and females develop from fertilized eggs and are diploid.

Many species in the genus alternate between an asexual generation and a sexual generation.

Species
The taxonomy of the genus Andricus is uncertain, with many of the approximately 375 species considered dubious. The following species are currently recognised in the genus Andricus:


 * Andricus aciculatus Beutenmuller, 1909
 * Andricus ahmeti Melika, Mutun & Dinç, 2014
 * Andricus albipes Hartig, 1840
 * Andricus albobalani Weld, 1926
 * Andricus alniensis Folliot, 1964
 * Andricus amblycerus (Giraud, 1859)
 * Andricus amenti Giraud, 1859
 * Andricus analis (Fonscolombe, 1832)
 * Andricus anatolicus Melika, Mutun & Dinç, 2014
 * Andricus archiboldi Melika & Abrahamson, 2021
 * Andricus aries (Girauld, 1859)
 * Andricus arslani Azmaz & Katılmış, 2022
 * Andricus assarehi Melika & Sadeghi, 2008
 * Andricus atkinsonae Melika, Stone, Sadeghi & Zargaran, 2008
 * Andricus bakrachus Melika, Mutun & Dinç, 2014
 * Andricus balanaspis (Weld, 1922)
 * Andricus balanella (Weld, 1957)
 * Andricus barriosi Medianero & Nieves-Aldrey, 2019
 * Andricus biconicus Weld, 1959
 * Andricus bimaculatus (Schenck, 1863)
 * Andricus brachycentrus (Thomson, 1877)
 * Andricus breviramuli Pujade-Villar, 2014
 * Andricus brunneus Fullaway, 1911 — clustered gall wasp
 * Andricus bulgaricus Vasileva-Symnalieva, 1978
 * Andricus burgundus Giraud, 1859
 * Andricus burnetti Dailey & Sprenger, 1983
 * Andricus caliciformis (Giraud, 1859)
 * Andricus californicus Ashmead, 1885
 * Andricus callidoma (Hartig, 1841)
 * Andricus capillatus (Weld, 1927)
 * Andricus caputmedusae (Hartig, 1843)
 * Andricus catalinensis Melika, Nicholls & Stone, 2021
 * Andricus catilla (Darboux & Houard, 1907)
 * Andricus cecconii Kieffer, 1901
 * Andricus chapmanii Melika & Abrahamson, 2021
 * Andricus chico Weld, 1957
 * Andricus chinquapin (Fitch, 1859) — small oak spindle gall wasp
 * Andricus chiricahuensis Melika, Nicholls & Stone, 2021
 * Andricus chodjaii Melika, 2008
 * Andricus chrysolepidicola Ashmead, 1896 — irregular spindle gall wasp
 * Andricus cinnamomeus Ashmead, 1887
 * Andricus clementinae (Giraud, 1859)
 * Andricus coconinoensis Melika, Nicholls & Stone, 2021
 * Andricus columbiensis Melika, Nicholls & Stone, 2021
 * Andricus comata Weld, 1956
 * Andricus confusus Lobato-Vila & Pujade-Villar, 2019
 * Andricus confertus McCracken & Egbert, 1922 — convoluted gall wasp
 * Andricus conglomeratus (Girauld, 1859)
 * Andricus conificus (Hartig, 1843)
 * Andricus cooki Melika, Nicholls & Stone, 2021
 * Andricus coortus Weld, 1947 — club gall wasp
 * Andricus coquilletti (Ashmead, 1897) - little oak apple gall wasp
 * Andricus coriariformis Melika, Challis & Stone, 2008
 * Andricus coriarius (Hartig, 1843)
 * Andricus coronatus (Girauld, 1859)
 * Andricus coronus Beutenmueller, 1907
 * Andricus corruptrix (Schlechtendal, 1870)
 * Andricus costaricensis Pujade-Villar & Melika, 2018
 * Andricus costatus Weld, 1944
 * Andricus crassicornis (Curtis, 1838)
 * Andricus crispator Tschek, 1871
 * Andricus cryptobius Wachtl, 1880
 * Andricus csokai Melika & Tavakoli, 2008
 * Andricus curtisii (Muller, 1870)
 * Andricus curvator (Hartig, 1840)
 * Andricus cydoniae Giraud, 1859
 * Andricus deciduatus Weld, 1926
 * Andricus dentimitratus (Rejtö, 1887)
 * Andricus dimorphus (Beutenmuller, 1913) — clustered midrib gall wasp
 * Andricus discalis (Weld, 1927)
 * Andricus exclusus (Ratzeburg, 1844)
 * Andricus fidelensis Kieffer, 1901
 * Andricus fitzpatricki Melika & Abrahamson, 2021
 * Andricus flavicornis Schenck, 1863
 * Andricus flavus Pujade-Villar, Wang, Guo & Chen, 2014
 * Andricus floridus Tavares, 1918
 * Andricus foecundatrix (Hartig, 1840)
 * Andricus flavohirtus Beutenmüller, 1913
 * Andricus foliaformis Weld, 1927
 * Andricus formosalis Weld, 1947
 * Andricus formosanus Tang & Melika, 2009
 * Andricus forni Pujade-Villar & Nicholls, 2020
 * Andricus frondeum (Weld, 1957)
 * Andricus fulviventris Schenck, 1863
 * Andricus furnaceus Kinsey, 1920
 * Andricus fuscicornis Hartig, 1841
 * Andricus galeatus (Girauld, 1859)
 * Andricus gallaecus Tavares, 1916
 * Andricus gallaetinctoriae (Olivier, 1791)
 * Andricus gallaeurnaeformis (Boyer de Fonscolombe, 1832)
 * Andricus gemmeus (Giraud, 1859)
 * Andricus gemmicola Kieffer, 1901
 * Andricus geniculatus (Dufour, 1864)
 * Andricus giardina Stefani, 1898
 * Andricus glandulae (Hartig, 1840)
 * Andricus glutinosus (Girauld, 1859)
 * Andricus gracilicornis (Kieffer, 1900)
 * Andricus grossulariae (Giraud, 1859)
 * Andricus hartigi (Hartig, 1843)
 * Andricus hastata (Kinsey, 1937)
 * Andricus hedwigia (Kustenmacher, 1894)
 * Andricus highlandensis Melika, Nicholls & Stone, 2021
 * Andricus hispanicus (Hartig, 1856)
 * Andricus howertoni Bassett, 1890
 * Andricus hungaricus (Hartig, 1843)
 * Andricus hyalinus Hartig, 1841
 * Andricus hystrix Kieffer, 1897
 * Andricus impropius Ballido & Pujade-Villar, 2003
 * Andricus incertus Bassett, 1900
 * Andricus indistinctus Bassett, 1890
 * Andricus infectorius (Hartig, 1843)
 * Andricus inflator Hartig, 1840
 * Andricus insanus ](Westwood, 1837)
 * Andricus istvani Melika, 2008
 * Andricus kashiwaphilus Abe, 1998
 * Andricus kollari (Hartig, 1843)
 * Andricus korlevici Kieffer, 1902
 * Andricus laevigatus Schenck, 1863
 * Andricus lasius (Ashmead, 1896) - hairy gall wasp
 * Andricus lateralis (Hartig, 1840)
 * Andricus legitimus Wiebes-Rijks, 1980
 * Andricus libani Melika, Challis & Stone, 2008
 * Andricus lignicolus (Hartig, 1840)
 * Andricus longipennis (Ashmead, 1887)
 * Andricus lucidus (Hartig, 1843)
 * Andricus luisieri Tavares, 1914
 * Andricus luteicornis Kieffer, 1899
 * Andricus malpighii (Adler, 1881)
 * Andricus mammadovi Azmaz & Katilmis, 2021
 * Andricus mamillaformis (Weld, 1926)
 * Andricus marmoreus Kinsey, 1920
 * Andricus maxwelli Bassett, 1890
 * Andricus mayri (Wachtl, 1881)
 * Andricus megatruncicolus Melika, 2008
 * Andricus melikai Pujade-Villar & Kwast, 2002
 * Andricus mellificus Melika, Nicholls & Stone, 2021
 * Andricus mendocinensis Weld, 1957
 * Andricus menkei Melika & Abrahamson, 2021
 * Andricus miriami Shachar, 2015
 * Andricus mitratus (Mayr, 1870)
 * Andricus mogollonensis Melika, Nicholls & Stone, 2021
 * Andricus montezumus Beutenmuller, 1913
 * Andricus moniliatus Hartig, 1840
 * Andricus moreae (Graeffe, 1905)
 * Andricus morula Shachar, Inbar & Dorchin, 2017
 * Andricus mukaigawae (Mukaigawa, 1913)
 * Andricus multiplicatus (Girauld, 1859)
 * Andricus murtfeldtae Ashmead, 1896
 * Andricus nichollsi Melika & Stone, 2021
 * Andricus niger (Fourcroy, 1785)
 * Andricus nigricens (Gillette, 1888)
 * Andricus nobrei Tavares, 1901
 * Andricus nodifex Kieffer, 1900
 * Andricus notholithocarpi Melika, Nicholls & Stone, 2018
 * Andricus occultatus (Weld, 1926)
 * Andricus octosporifex (Schrank, 1802)
 * Andricus opertus (Weld, 1926) — fimbriate gall wasp
 * Andricus ostrea (Hartig, 1840)
 * Andricus pallidicornis (Curtis, 1838)
 * Andricus pallipes (Schenck, 1863)
 * Andricus panteli (Kieffer, 1896)
 * Andricus paradoxus (Radoszkowski, 1866)
 * Andricus partali Pujade-Villar, Cuesta-Porta & Hanson, 2022
 * Andricus pedicellatus Kinsey, 1922 — hair stalk gall wasp
 * Andricus peredurus (Kinsey, 1920)
 * Andricus perfulvum (Weld, 1952)
 * Andricus perlentus (Kinsey, 1937)
 * Andricus petioli Hartig, 1843
 * Andricus pictus (Hartig, 1856)
 * Andricus pilula (Bassett, 1890)
 * Andricus pilularis Weld, 1952
 * Andricus pisiformis Beutenmueller, 1911
 * Andricus polycerus (Girauld, 1859)
 * Andricus prescotti Weld, 1952
 * Andricus projectus Weld, 1952
 * Andricus pseudoaries Melika, Stone & Sadegh, 2008
 * Andricus pseudocurvator Tang & Melika, 2011
 * Andricus pseudoinflator Tavares, 1901
 * Andricus pujadevillari Melika, Stone, Sadeghi, Atkinson & Zargaran, 2008
 * Andricus quadrilineatus (Hartig, 1840)
 * Andricus quercuscalicis (Burgsdorf, 1783) — knopper gall
 * Andricus quercuscalifornicus (Bassett, 1881) — California gall wasp
 * Andricus quercuscorticis (Linnaeus, 1761)
 * Andricus quercusfoliatus (Ashmead, 1881) — leafy oak gall wasp
 * Andricus quercusformosus (Bassett, 1864)
 * Andricus quercusfrondosus (Bassett, 1865)
 * Andricus quercusinferus (Linnaeus, 1767)
 * Andricus quercuslaurinus Melika and Pujade-Villar, 2009
 * Andricus quercuspetioli (Linnaeus, 1758)
 * Andricus quercuspetiolicola Bassett, 1863 — oak petiole gall wasp
 * Andricus quercuspyramidalis (Landois, 1895)
 * Andricus quercusradicis (Fabricius, 1798)
 * Andricus quercusramuli (Linnaeus, 1761)
 * Andricus quercussingularis Bassett, 1863 — small oak apple gall wasp
 * Andricus quercustozae (Bosc, 1792)
 * Andricus quercusstrobilanus (Osten-Sacken, 1862) — lobed oak gall wasp
 * Andricus quercusutriculus (Bassett, 1881)
 * Andricus reticulatus Bassett, 1890
 * Andricus rhizoxenus (Ashmead, 1896)
 * Andricus rhyzomae (Hartig, 1843)
 * Andricus robustus Weld, 1926
 * Andricus rotundula (Weld, 1952)
 * Andricus rubripes (Thomson, 1877)
 * Andricus ruficornis (Schenck, 1863)
 * Andricus rufipes (Fabricius, 1804)
 * Andricus rufiventris Schenck, 1863
 * Andricus rufus (Thomson, 1877)
 * Andricus rugatus Weld, 1926
 * Andricus ruginosus Bassett, 1890
 * Andricus sadeghii Melika, Stone, Atkinson & Aligolizade, 2008
 * Andricus schencki Dalla Torre & Kieffer, 1910
 * Andricus schickae Nicholls, Melika & Stone, 2021
 * Andricus schoenroggei Melika & Stone, 2008
 * Andricus schroeckingeri Wachtl, 1876
 * Andricus scutella Weld, 1930
 * Andricus seckendorffi (Wachtl, 1879)
 * Andricus seminationis (Giraud, 1859)
 * Andricus serotinus (Giraud, 1859)
 * Andricus serricornis (Kinsey, 1922)
 * Andricus sessilum Weld, 1926
 * Andricus shuhuti Melika, Mutun & Dinç, 2013
 * Andricus sieboldi (Hartig, 1843)
 * Andricus singularis Mayr, 1870
 * Andricus solitarius (Fonscolombe, 1832)
 * Andricus songshui Tang & Melika, 2011
 * Andricus sphaericus Pujade-Villar, 2016
 * Andricus spicatus (Bassett, 1900)
 * Andricus splendens Weld, 1919
 * Andricus stefanii (Kieffer, 1897)
 * Andricus stellaris Weld, 1926 — sunburst gall wasp
 * Andricus stellulus (Burnett, 1974) — stellar gall wasp
 * Andricus sternlichti Bellido, Pujade-Villar & Melika, 2003
 * Andricus stonei Melika, Tavakoli & Sadeghi, 2014
 * Andricus stramineus Weld, 1944
 * Andricus stropus Ashmead, 1887
 * Andricus strues (Kinsey, 1938)
 * Andricus subterranea (Giraud, 1859)
 * Andricus sufflator Mayr, 1882
 * Andricus sulfureus Weld, 1926
 * Andricus superfetationis (Giraud, 1859)
 * Andricus surculi (Schrank, 1781)
 * Andricus tavaresi Kieffer, 1904
 * Andricus testaceipes (Hartig, 1840)
 * Andricus testaceus (Gmelin, 1790)
 * Andricus texanus Beutenmüller, 1909
 * Andricus theophrasteus (Trotter, 1902)
 * Andricus tomentosus (Trotter, 1901)
 * Andricus torreyaensis Melika & Abrahamson, 2021
 * Andricus toumeyi Weld, 1926
 * Andricus trotteri Kieffer, 1898
 * Andricus truncicola (Girauld, 1859)
 * Andricus tubalis Weld, 1926
 * Andricus tubularius Weld, 1926
 * Andricus tumefaciens Pujade-Villar & Paretas-Martínez, 2014
 * Andricus turcicus Melika, Mutun & Dinç, 2014
 * Andricus turionum (Hartig, 1840)
 * Andricus vacciniifoliae (Ashmead, 1896) — oak apple wasp
 * Andricus villosulus (Gravenhorst, 1807)
 * Andricus verensis Weld, 1955
 * Andricus vindobonensis Müllner, 1901
 * Andricus weldi (Beutenmüller, 1918)
 * Andricus wheeleri Beutenmueller, 1907
 * Andricus williami Melika, Nicholls & Stone, 2021
 * Andricus wiltzae Fullaway, 1911
 * Andricus zappellai Kieffer, 1901

Previous species
These species were previously grouped in Andricus.

Currently in Druon:
 * Druon flocculentum Lyon
 * Druon fullawayi Beutenmüller, 1914
 * Druon ignotum Bassett
 * Druon linaria Kinsey
 * Druon pattoni (Bassett, 1881)
 * Druon protagion Kinsey
 * Druon quercusflocci (Walsh, 1864)
 * Druon quercuslanigerum Ashmead

Currently in Feron:


 * Feron albicomus (Weld, 1952)
 * Feron amphorus (Weld, 1926)
 * Feron apiarium (Weld, 1944)
 * Feron atrimentum Kinsey, 1922) — striped volcano gall wasp
 * Feron bakkeri (Lyon, 1984)
 * Feron caepula (Weld, 1926)
 * Feron californicum (Beutenmueller, 1911)
 * Feron clarkei (Bassett, 1890)
 * Feron crystallinum (Bassett, 1900) — crystalline gall wasp
 * Feron cylindratum (Kinsey, 1937)
 * Feron discularis (Weld, 1926)
 * Feron gigas (Kinsey, 1922) —saucer gall wasp
 * Feron kingi (Bassett, 1900) — red cone gall wasp
 * Feron parmula (Bassett, 1900) — disc gall wasp
 * Feron pattersonae Fullaway, 1911 — plate gall wasp
 * Feron tecturnarum Kinsey, 1920
 * Feron tubifaciens Weld, 1926