Bothriembryontidae

Bothriembryontidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Orthalicoidea.

Taxonomy
This family has been named Placostylidae and had no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Breure et al. (2010) moved Prestonella and Bothriembryon to Placostylidae.

Breure & Romero (2012) confirmed previous results from 2010 and they renamed Placostylidae to Bothriembryontidae.


 * subfamily Bothriembryontinae Iredale, 1937
 * subfamily Placostylinae Pilsbry, 1946 - for species from New Zealand and Melanesia

Prestonellinae was formally described as a new subfamily within Bothriembryontidae in 2016. Therefore three subfamilies are recognized in Bothriembryontidae:
 * subfamily Bothriembryontinae Iredale, 1937
 * subfamily Placostylinae Pilsbry, 1946
 * Prestonellinae van Bruggen, Herbert & Breure, 2016

Genera
Genera within the family Bothriembryontidae include:

Bothriembryontinae
 * Bothriembryon - the type genus of the family Bothriembryontidae

Placostylinae
 * Archaeostylus Brook & Hayward, 2022
 * Aspastus Albers, 1850
 * Basileostylus F. Haas, 1935
 * Callistocharis Pilsbry, 1900
 * Diplomorpha Ancey, 1884
 * Eumecostylus Martens in Albers, 1860
 * Euplacostylus Crosse, 1875
 * Leucocharis Pilsbry, 1900
 * Placocharis Pilsbry, 1900
 * Placostylus Beck, 1837 - the type genus of the (sub)family Placostylidae
 * Poecilocharis Kobelt, 1891
 * Quiros Solem, 1959
 * Santacharis Iredale, 1927

Prestonellinae
 * Discoleus Breure, 1978
 * Plectostylus Beck, 1837
 * Prestonella Connolly, 1929 - type genus of the subfamily Prestonellinae

A cladogram based on nuclear ITS2, 28S rRNA and histone 3 (H3) and sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of Bothriembryontidae by Breure & Romero (2012):

Distribution
Bothriembryontidae have Gondwanan distribution.