Bryaxis (beetle)

Bryaxis is a genus of ant-loving beetles in the family Staphylinidae. There are more than 380 described species in Bryaxis.

Bryaxis species
These 385 species are members of the genus Bryaxis.


 * Bryaxis abastumanus (Reitter, 1881)
 * Bryaxis abkhasicus Besuchet & Kurbatov, 2007
 * Bryaxis acernus Castellini, 1978
 * Bryaxis adjaricus Besuchet & Kurbatov, 2007
 * Bryaxis adumbratus Besuchet & Kurbatov, 2007
 * Bryaxis aelistae (Reitter, 1882)
 * Bryaxis affinis (Sharp, 1883)
 * Bryaxis alayai Löbl & Kurbatov, 1996
 * Bryaxis albanicus (Apfelbeck, 1907)
 * Bryaxis alesi Löbl & Kurbatov, 1996
 * Bryaxis alishanus Löbl & Kurbatov, 1996
 * Bryaxis alpestris (Dodero, 1900)
 * Bryaxis altivagus Besuchet, 1962
 * Bryaxis amasiae (Reitter, 1890)
 * Bryaxis ami Löbl & Kurbatov, 1996
 * Bryaxis ammon (Saulcy, 1878)
 * Bryaxis amurensis Kurbatov, 1985
 * Bryaxis anatolicus (Saulcy, 1878)
 * Bryaxis andalusiacus (Jeannel, 1956)
 * Bryaxis appendiculatus (Reitter, 1885)
 * Bryaxis aragonensis (Franz, 1955)
 * Bryaxis argiolus (Reitter, 1888)
 * Bryaxis argodi (Croissandeau, 1885)
 * Bryaxis argus (Kraatz, 1863)
 * Bryaxis armeniacus Brachat, 2019
 * Bryaxis armipes (Reitter, 1881)
 * Bryaxis arnoldii Besuchet, 1961
 * Bryaxis artvinensis Besuchet & Kurbatov, 2007
 * Bryaxis asciicornis Kurbatov, 1985
 * Bryaxis assingi Besuchet & Kurbatov, 2007
 * Bryaxis asturiensis (Reitter, 1880)
 * Bryaxis atilla (Saulcy, 1878)
 * Bryaxis atlanticides Newton, 2015
 * Bryaxis auritides Newton, 2015
 * Bryaxis badius Besuchet, 1961
 * Bryaxis bajulus (Hampe, C., 1863)
 * Bryaxis balabanus Besuchet & Kurbatov, 2007
 * Bryaxis balneator Besuchet & Kurbatov, 2007
 * Bryaxis bedeki Bekchiev & Hlaváč, 2016
 * Bryaxis bellax Löbl & Kurbatov, 1996
 * Bryaxis bergamascus (Stolz, 1917)
 * Bryaxis bericus Pace, 1974
 * Bryaxis beroni Karaman, 1969
 * Bryaxis biokovensis Bekchiev & Hlaváč, 2016
 * Bryaxis blacensis (Karaman, 1954)
 * Bryaxis bodemeyeri (Reitter, 1903)
 * Bryaxis bojanensis (Apfelbeck, 1907)
 * Bryaxis borckensis Besuchet & Kurbatov, 2007
 * Bryaxis bosnicus (Ganglbauer, 1895)
 * Bryaxis bothrophorus (Stolz, 1917)
 * Bryaxis brachati Besuchet, 1980
 * Bryaxis brescianus Daffner, 1986
 * Bryaxis brusinae (Reitter, 1880)
 * Bryaxis buddha Kurbatov & Löbl, 1995
 * Bryaxis bulbifer (Reichenbach, 1816)
 * Bryaxis bushido Löbl, Kurbatov & Nomura, 1998
 * Bryaxis callipus (Apfelbeck, 1906)
 * Bryaxis carinula (Rey, 1888)
 * Bryaxis carpathicus (Saulcy, 1875)
 * Bryaxis casalei Poggi, 1992
 * Bryaxis castanoptera Motschulsky, 1851
 * Bryaxis catalanus (Jeannel, 1950)
 * Bryaxis cateniger (Krauss, 1899)
 * Bryaxis cebennicus (Jeannel, 1950)
 * Bryaxis cephalotes (Motschulsky, 1845)
 * Bryaxis cepliki Hlaváč, 2009
 * Bryaxis cetinjensis (Apfelbeck, 1906)
 * Bryaxis chaudoirii (Chaudoir, 1845)
 * Bryaxis chevrolati (Aubé, 1833)
 * Bryaxis chiarae Monguzzi, 2016
 * Bryaxis chilimontanus Nomura & Chang Eon Lee, 1993
 * Bryaxis chobautianus Löbl, 1998
 * Bryaxis claudioi Daffner, 1992
 * Bryaxis clavicornis (Panzer, 1805)
 * Bryaxis clavipes (Motschulsky, 1851)
 * Bryaxis cocles (Saulcy, 1863)
 * Bryaxis coelestis Löbl & Kurbatov, 1996
 * Bryaxis coiffaiti Jeannel, 1958
 * Bryaxis collaris (Baudi di Selve, 1859)
 * Bryaxis colledanii Daffner, 1984
 * Bryaxis comita (Rambousek, 1909)
 * Bryaxis convexus (Kiesenwetter, 1858)
 * Bryaxis corcyreus (Reitter, 1884)
 * Bryaxis cornutus (Machulka, 1938)
 * Bryaxis corsus Besuchet, 1999
 * Bryaxis crassicornides Newton, 2017
 * Bryaxis crassicornis (Motschulsky, 1835)
 * Bryaxis credibilis Besuchet & Kurbatov, 2007
 * Bryaxis crepsensis (Müller, G., 1947)
 * Bryaxis crotchi (Sharp, 1874)
 * Bryaxis curtisii (Leach, 1817)
 * Bryaxis cythereias (Pic, 1903)
 * Bryaxis dalmatinus (Reitter, 1881)
 * Bryaxis dichrous (Reitter, 1882)
 * Bryaxis difficilis (Reitter, 1884)
 * Bryaxis disciger (Roubal, 1910)
 * Bryaxis distinguendus Besuchet, 1961
 * Bryaxis diversicornis (Raffray, 1873)
 * Bryaxis dolosus Poggi & Sardu, 2013
 * Bryaxis effeminatus Besuchet, 1983
 * Bryaxis egens Besuchet & Kurbatov, 2007
 * Bryaxis emeicus Kurbatov & Löbl, 1995
 * Bryaxis emendatus Besuchet & Kurbatov, 2007
 * Bryaxis emilianus (Stolz, 1926)
 * Bryaxis erichsonii (Kiesenwetter, 1849)
 * Bryaxis eurus Karaman, 1967
 * Bryaxis euryscapus Besuchet & Kurbatov, 2007
 * Bryaxis extremalis Kurbatov, 1990
 * Bryaxis fagei (Jeannel, 1926)
 * Bryaxis fauconneti (Fauvel, 1886)
 * Bryaxis femoratus (Aubé, 1844)
 * Bryaxis festivus Besuchet, 1964
 * Bryaxis fictor Kurbatov & Löbl, 1998
 * Bryaxis focarilei Besuchet, 1980
 * Bryaxis fossicornis (Rambousek, 1916)
 * Bryaxis frivaldszkyi (Reitter, 1887)
 * Bryaxis fronticornis Bekchiev, 2013
 * Bryaxis frustratus Besuchet, 1983
 * Bryaxis gallicus (Reitter, 1887)
 * Bryaxis ganglbaueri (Stolz, 1917)
 * Bryaxis gemellus Besuchet & Kurbatov, 2007
 * Bryaxis ghilarovi Besuchet, 1961
 * Bryaxis giraffa (Reitter, 1880)
 * Bryaxis goliath (Jeannel, 1922)
 * Bryaxis gracilipalpis Jeannel, 1958
 * Bryaxis grandicollis Jeannel, 1958
 * Bryaxis grilati (Reitter, 1886)
 * Bryaxis grouvellei (Reitter, 1882)
 * Bryaxis hakka Löbl & Kurbatov, 1996
 * Bryaxis halbherri (Reitter, 1885)
 * Bryaxis hallamontanus Nomura & Chang Eon Lee, 1992
 * Bryaxis harmandi Raffray, 1909
 * Bryaxis heian Löbl, Kurbatov & Nomura, 1998
 * Bryaxis heilongjiangensis Li, Jingke, 1993
 * Bryaxis heliophobus (Apfelbeck, 1906)
 * Bryaxis herculinus Besuchet, 1962
 * Bryaxis heydeni (Reitter, 1880)
 * Bryaxis hiranoi Taru & Nomura, 2021
 * Bryaxis hisamatsui Löbl, Kurbatov & Nomura, 1998
 * Bryaxis hokkiensis Löbl & Kurbatov, 1996
 * Bryaxis hoko Löbl, Kurbatov & Nomura, 1998
 * Bryaxis holciki Löbl, 1964
 * Bryaxis humilis Raffray, 1909
 * Bryaxis hypocritus Plavilstschikov, 1940
 * Bryaxis ibericus (Saulcy, 1870)
 * Bryaxis immodicus Besuchet & Kurbatov, 2007
 * Bryaxis indistincta Motschulsky, 1851
 * Bryaxis insularis (Holdhaus, 1905)
 * Bryaxis ipsimus Besuchet & Kurbatov, 2007
 * Bryaxis iriomotensis Löbl, Kurbatov & Nomura, 1998
 * Bryaxis islamitus (Reitter, 1885)
 * Bryaxis issensis (Müller, J., 1909)
 * Bryaxis italicus (Baudi di Selve, 1870)
 * Bryaxis japonicus (Sharp, 1874)
 * Bryaxis joffrei (Sainte-Claire Deville, 1928)
 * Bryaxis jomon Löbl, Kurbatov & Nomura, 1998
 * Bryaxis jucundus Besuchet, 1961
 * Bryaxis judaeus (Pic, 1900)
 * Bryaxis judicariensis (Dodero, 1919)
 * Bryaxis kahleni Pace, 1978
 * Bryaxis kamakura Löbl, Kurbatov & Nomura, 1998
 * Bryaxis karamanae Besuchet, 1958
 * Bryaxis karaormani (Karaman, 1954)
 * Bryaxis karate Löbl, Kurbatov & Nomura, 1998
 * Bryaxis katana Löbl, Kurbatov & Nomura, 1998
 * Bryaxis khnzoriani Besuchet, 1964
 * Bryaxis kimi Kurbatov & Löbl, 1998
 * Bryaxis kimjongkuki Nomura & Chang Eon Lee, 1993
 * Bryaxis klimeschi (Blattný, W. & C. Blattný, 1914)
 * Bryaxis kofun Löbl, Kurbatov & Nomura, 1998
 * Bryaxis koltzei (Reitter, 1887)
 * Bryaxis konecznii (Machulka, 1938)
 * Bryaxis kovali Besuchet & Kurbatov, 2007
 * Bryaxis krilei Hlaváč, 2008
 * Bryaxis kruegeri (Machulka, 1932)
 * Bryaxis kumensis Daffner, 1992
 * Bryaxis kumgangsanensis Löbl, 2000
 * Bryaxis kurnakovi Besuchet, 1960
 * Bryaxis kuzmini Besuchet & Kurbatov, 2007
 * Bryaxis laevipennis Besuchet & Kurbatov, 2007
 * Bryaxis lagari (Halbherr, 1890)
 * Bryaxis lanai Poggi, 2019
 * Bryaxis lata Motschulsky, 1851
 * Bryaxis lazistanicus Besuchet & Kurbatov, 2007
 * Bryaxis lederi (Reitter, 1888)
 * Bryaxis leechanyoungi Nomura & Chang Eon Lee, 1993
 * Bryaxis leleupi (Jeannel, 1952)
 * Bryaxis leonhardi (Reitter, 1902)
 * Bryaxis lesbius Brachat, 2016
 * Bryaxis lessinicus Pace, 1974
 * Bryaxis libanicola (Sahlberg, J., 1908)
 * Bryaxis liguricus (Dodero, 1919)
 * Bryaxis litoralis Besuchet & Kurbatov, 2007
 * Bryaxis lokayi (Machulka, 1927)
 * Bryaxis longifrons Jeannel, 1952
 * Bryaxis longifronsides Newton, 2015
 * Bryaxis longipalpis (Motschulsky, 1835)
 * Bryaxis longulus (Kiesenwetter, 1849)
 * Bryaxis lurensis Besuchet, 2002
 * Bryaxis lusitanicus (Saulcy, 1870)
 * Bryaxis macedonicus (Karaman, 1953)
 * Bryaxis macheonensis Nomura & Chang Eon Lee, 1993
 * Bryaxis mahunkai Löbl, 1975
 * Bryaxis mancinii (Dodero, 1919)
 * Bryaxis mandarinus Löbl & Kurbatov, 1996
 * Bryaxis marinae Sabella, 1989
 * Bryaxis mayumi Löbl, Kurbatov & Nomura, 1998
 * Bryaxis meghruicus Brachat, 2019
 * Bryaxis meledanus Agazzi, 1964
 * Bryaxis melinensis (Reitter, 1881)
 * Bryaxis mendax Kurbatov & Löbl, 1998
 * Bryaxis merkli Löbl, 2000
 * Bryaxis meskischesianus Meggiolaro, 1966
 * Bryaxis meybohmianus Brachat, 2019
 * Bryaxis mirei (Jeannel, 1956)
 * Bryaxis mirificus Besuchet, 1983
 * Bryaxis moczarskii (Winkler, A., 1911)
 * Bryaxis mohamedis (Reitter, 1903)
 * Bryaxis monguzzii Besuchet, 1980
 * Bryaxis monstrosetibialis (Stolz, 1923)
 * Bryaxis monticola (Jeannel, 1950)
 * Bryaxis montivagus (Reitter, 1885)
 * Bryaxis mosorensis (Karaman, 1953)
 * Bryaxis mulsanti Kiesenwetter, 1850
 * Bryaxis multiplex Besuchet & Kurbatov, 2007
 * Bryaxis muscorum (Kiesenwetter, 1849)
 * Bryaxis myops Besuchet & Kurbatov, 2007
 * Bryaxis naginata Löbl, Kurbatov & Nomura, 1998
 * Bryaxis nakeralae (Reitter, 1885)
 * Bryaxis nametkini Kurbatov & Löbl, 1998
 * Bryaxis nebrodensis Besuchet, 1980
 * Bryaxis nemilensis (Reitter, 1885)
 * Bryaxis nigriceps (Leach, 1826)
 * Bryaxis nigripennis (Aubé, 1844)
 * Bryaxis nigritus Löbl, 1998
 * Bryaxis nitidulus Besuchet, 1961
 * Bryaxis nivarius Besuchet & Kurbatov, 2007
 * Bryaxis nodicornis (Aubé, 1833)
 * Bryaxis nogodanensis Nomura & Chang Eon Lee, 1993
 * Bryaxis noricus (Schubert, F., 1958)
 * Bryaxis normandi (Jeannel, 1950)
 * Bryaxis obenbergeri (Machulka, 1938)
 * Bryaxis obventicius Besuchet & Kurbatov, 2007
 * Bryaxis odontogena (Dodero, 1919)
 * Bryaxis oertzeni (Reitter, 1882)
 * Bryaxis oraniensis (Pic, 1896)
 * Bryaxis orcinus Besuchet & Kurbatov, 2007
 * Bryaxis oreophilus (Meixner, 1912)
 * Bryaxis orousseti Secq, B. & M. Secq, 1991
 * Bryaxis osellai Besuchet & Kurbatov, 2007
 * Bryaxis oseticus Besuchet & Kurbatov, 2007
 * Bryaxis ossaeus Besuchet, 2008
 * Bryaxis pachyscelis Besuchet & Kurbatov, 2007
 * Bryaxis paganettii (Blattný, W. & C. Blattný, 1914)
 * Bryaxis paiwan Löbl & Kurbatov, 1996
 * Bryaxis pallidior (Pic, 1901)
 * Bryaxis panda Kurbatov & Löbl, 1995
 * Bryaxis pandellei (Saulcy, 1863)
 * Bryaxis pangeonicus Brachat, 2019
 * Bryaxis pastoralis (Peyerimhoff, 1901)
 * Bryaxis pavani (Tamanini, 1940)
 * Bryaxis pawlowskii Löbl, 1974
 * Bryaxis peckorum Löbl, Kurbatov & Nomura, 1998
 * Bryaxis pedator (Reitter, 1882)
 * Bryaxis pedemontanus Besuchet, 1958
 * Bryaxis pedestris (Motschulsky, 1845)
 * Bryaxis peloponnesius (Reitter, 1884)
 * Bryaxis peninsulanus Nomura & Chang Eon Lee, 1993
 * Bryaxis peninsularis (Saulcy, 1870)
 * Bryaxis pentagonoceras (Stolz, 1917)
 * Bryaxis persicoi (Rasetti, F., 1939)
 * Bryaxis pescaroloi Poggi, 1984
 * Bryaxis petri Raffray, 1915
 * Bryaxis pictetii (Tournier, 1859)
 * Bryaxis pilifera Motschulsky, 1851
 * Bryaxis pingtungensis Löbl & Kurbatov, 1996
 * Bryaxis pinkeri (Stolz, 1917)
 * Bryaxis planifer Löbl & Kurbatov, 1996
 * Bryaxis platalea Löbl, Kurbatov & Nomura, 1998
 * Bryaxis pletnevi Kurbatov & Löbl, 1998
 * Bryaxis polemon Besuchet & Kurbatov, 2007
 * Bryaxis ponticus Besuchet & Kurbatov, 2007
 * Bryaxis porsenna Reitter, 1882
 * Bryaxis portalegrensis (Schaufuss, L. W., 1882)
 * Bryaxis procerus (Gredler, 1873)
 * Bryaxis propinquus Besuchet & Kurbatov, 2007
 * Bryaxis pulchrotibialis Besuchet & Kurbatov, 2007
 * Bryaxis pumilus Bekchiev & Brachat, 2015
 * Bryaxis puncticollis (Denny, 1825)
 * Bryaxis pygmaeides Newton, 2015
 * Bryaxis pyrenaeus (Saulcy, 1863)
 * Bryaxis rambouseki (Matcha, 1916)
 * Bryaxis raptor Löbl & Kurbatov, 1996
 * Bryaxis redemptus (Dodero, 1919)
 * Bryaxis reissi (Machulka, 1933)
 * Bryaxis reitteri (Saulcy, 1875)
 * Bryaxis reversus (Sharp, 1883)
 * Bryaxis rhinophorus (Blattný, W. & C. Blattný, 1914)
 * Bryaxis rifensis Besuchet, 1962
 * Bryaxis rivularis Besuchet & Kurbatov, 2007
 * Bryaxis rossii Castellini, 1986
 * Bryaxis rosti (Reitter, 1894)
 * Bryaxis rostratus (Motschulsky, 1845)
 * Bryaxis roubalianus Löbl, 1961
 * Bryaxis roumaniae Raffray, 1904
 * Bryaxis rousi Besuchet & Kurbatov, 2007
 * Bryaxis rugosicollis (Fiori, 1905)
 * Bryaxis ruidus Kurbatov & Löbl, 1998
 * Bryaxis ruthenus (Saulcy, 1877)
 * Bryaxis sacer Kurbatov & Löbl, 1995
 * Bryaxis sacrificus Kurbatov & Löbl, 1995
 * Bryaxis samniticus (Blattný, W. & C. Blattný, 1914)
 * Bryaxis samothracicus Brachat, 2019
 * Bryaxis samurai Löbl, Kurbatov & Nomura, 1998
 * Bryaxis sarplaninensis (Karaman, 1953)
 * Bryaxis sauteri Raffray, 1909
 * Bryaxis sawadai Löbl, Kurbatov & Nomura, 1998
 * Bryaxis scapularis (Reitter, 1881)
 * Bryaxis schuelkei Besuchet & Kurbatov, 2007
 * Bryaxis schusteri Neuhäuser-Happe, 1999
 * Bryaxis scrutandus Besuchet & Kurbatov, 2007
 * Bryaxis sculpticornis (Guillebeau, 1891)
 * Bryaxis sculptifrons (Reitter, 1880)
 * Bryaxis seductus Besuchet & Kurbatov, 2007
 * Bryaxis serripes (Fauvel, 1891)
 * Bryaxis sichotensis Kurbatov, 1994
 * Bryaxis sichuanus Kurbatov & Löbl, 1995
 * Bryaxis siculus (Fiori, 1913)
 * Bryaxis silvicolides Newton, 2015
 * Bryaxis simoni (Reitter, 1880)
 * Bryaxis smetanai Löbl, 1964
 * Bryaxis solidus (Reitter, 1881)
 * Bryaxis sparcepunctatus Jeannel, 1958
 * Bryaxis splendidus (Croissandeau, 1891)
 * Bryaxis steindachneri (Reitter, 1881)
 * Bryaxis stolzi (Machulka, 1932)
 * Bryaxis sturanyi (Apfelbeck, 1906)
 * Bryaxis subdentatus (Dodero, 1919)
 * Bryaxis subseriatus (Weise, J., 1877)
 * Bryaxis subsolidus (Reitter, 1902)
 * Bryaxis sumo Löbl, Kurbatov & Nomura, 1998
 * Bryaxis swaneticus (Reitter, 1885)
 * Bryaxis taiwanus Löbl & Kurbatov, 1996
 * Bryaxis tanto Löbl, Kurbatov & Nomura, 1998
 * Bryaxis taradakensis Löbl, Kurbatov & Nomura, 1998
 * Bryaxis tarokanus Löbl & Kurbatov, 1996
 * Bryaxis temporalis Besuchet & Kurbatov, 2007
 * Bryaxis tendensis Besuchet, 2002
 * Bryaxis tenuicornides Newton, 2015
 * Bryaxis testatus Kurbatov, 1994
 * Bryaxis tetralobus Löbl, Kurbatov & Nomura, 1998
 * Bryaxis theanus (Reitter, 1894)
 * Bryaxis tienmushanus Kurbatov & Löbl, 1995
 * Bryaxis tingitanus Besuchet, 1962
 * Bryaxis tithonus (Apfelbeck, 1906)
 * Bryaxis transitorius Besuchet & Kurbatov, 2007
 * Bryaxis transsilvanicus (Ganglbauer, 1897)
 * Bryaxis trezzii Monguzzi, 2016
 * Bryaxis trichosus Jeannel, 1958
 * Bryaxis trigonoceras (Holdhaus, 1904)
 * Bryaxis troglocerus (Saulcy, 1870)
 * Bryaxis troglodytes (Fiori, 1900)
 * Bryaxis troglophilus Agazzi, 1961
 * Bryaxis tscherkessicus (Reitter, 1888)
 * Bryaxis tuberculiceps Nonveiller, Pavićević & Besuchet, 2003
 * Bryaxis tychobythinoides Löbl & Kurbatov, 1996
 * Bryaxis ullrichii (Motschulsky, 1851)
 * Bryaxis unicus Löbl & Kurbatov, 1996
 * Bryaxis ursus (Reitter, 1882)
 * Bryaxis ussuriensis Löbl, 1964
 * Bryaxis valentulus Kurbatov & Löbl, 1998
 * Bryaxis validicornides Newton, 2015
 * Bryaxis verrucipalpis (Apfelbeck, 1907)
 * Bryaxis vicinus (Dodero, 1919)
 * Bryaxis viertli (Reitter, 1882)
 * Bryaxis viti Besuchet & Kurbatov, 2007
 * Bryaxis vlastae Roubal, 1913
 * Bryaxis wankai (Reitter, 1915)
 * Bryaxis weisei (Saulcy, 1875)
 * Bryaxis witzgalli Daffner, 1982
 * Bryaxis wolongensis Kurbatov & Löbl, 1995
 * Bryaxis yari Löbl, Kurbatov & Nomura, 1998
 * Bryaxis ypsilon Besuchet & Kurbatov, 2007
 * Bryaxis yushanensis Löbl & Kurbatov, 1996
 * Bryaxis zubroides Bekchiev, 2022