Bulimulidae

Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.

Distribution
Distribution of species in the family Bulimulidae includes Ecuador (9 genera) and other South American countries. Some species also occur in North America.

Fossil record
The family's oldest fossil record dates from the late Cretaceous of Brazil (Itaboraí Basin).

Anatomy
Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).

Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4–1/6 total phallus length).

Taxonomy
Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.

2005 taxonomy
Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).

The subfamily Bulimulinae included 3 tribes:
 * Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
 * Tribe Odontostomini Pilsbry & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
 * Tribe Simpulopsini Schileyko, 1999

2010 taxonomy
Breure et al. (2010) moved Bothriembryon to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae.

2012 taxonomy
Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae. There are three subfamilies within Bulimulidae:


 * subfamily Bulimulinae Tryon, 1867
 * subfamily Peltellinae Gray, 1855
 * subfamily Bostrycinae Breure, 2012

Genera
Genera in the family Bulimulidae include:
 * subfamily Bulimulinae Tryon, 1887 : (synonym: Berendtiinae P. Fischer & Crosse, 1872 )
 * Anctus E. von Martens, 1860
 * Auris Spix, 1827
 * Berendtia Crosse & P. Fischer, 1869
 * Bocourtia Rochebrune, 1882
 * Bulimulus Leach, 1814 - type genus of the subfamily Bulimulinae
 * Cochlorina Jan, 1830
 * Graptostracus Pilsbry, 1939
 * Kora Simone, 2012
 * Lopesianus Weyrauch, 1958
 * Naesiotus Albers, 1850
 * Neopetraeus E. von Martens, 1885
 * Newboldius Pilsbry, 1932
 * Otostomus H. Beck, 1837
 * Oxychona Mörch, 1852
 * Protoglyptus Pilsbry, 1897
 * Pseudoxychona Pilsbry, 1930
 * Rabdotus Albers, 1850
 * Scutalus Albers, 1850
 * Spartocentrum Dall, 1895
 * Sphaeroconcha Breure, 1978
 * Stenostylus Pilsbry, 1898
 * Suniellus Breure, 1978
 * subfamily Peltellinae
 * Antidrymaeus Germain, 1907
 * Drymaeus Albers, 1850
 * Peltella Gray, 1855 - type genus of the subfamily Peltellinae
 * Mesembrinus Albers, 1850


 * subfamily Bostrycinae
 * Bostryx Troschel, 1847 sensu lato - type genus of the subfamily Bostrycinae


 * Unassigned within Bulimulidae
 * Bulimus Bruguière, 1789 (temporary name)
 * Itaborahia Maury, 1935
 * † Oreoconus D. W. Taylor in McKenna et al., 1962
 * † Palaeobulimulus Parodiz, 1949
 * Stapafurdius Simone, 2021
 * † Tocobaga Auffenberg, Slapcinsky & Portell, 2015

Synonyms

 * Adzharia P. Hesse, 1933 : synonym of Bulimulus Leach, 1814 (junior synonym)
 * Antidrymaeus Germain, 1907 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
 * Atahualpa Strebel, 1910 : synonym of Thaumastus E. von Martens, 1860
 * Ataxellus Dall, 1912 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented asBostryx Troschel, 1847
 * Ataxus Albers, 1850 : synonym of Bostryx Troschel, 1847
 * Bilamelliferus Weyrauch, 1958 : synonym of Bostryx Troschel, 1847 (junior synonymy)
 * Cochlogena Férussac, 1821 : synonym of Bulimulus Leach, 1814 (junior synonym)
 * Diaphanomormus Weyrauch, 1964 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
 * Elatibostryx Weyrauch, 1958 : synonym of Bostryx Troschel, 1847
 * Floreziellus Weyrauch, 1967 : synonym of Bostryx Troschel, 1847 (junior synonymy)
 * Geoceras Pilsbry, 1896 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
 * Globulinus Crosse & P. Fischer, 1875 : synonym of Rabdotus Albers, 1850
 * Goniognathmus Crosse & P. Fischer, 1875 : synonym of Drymaeus Albers, 1850 (junior synonym)
 * Hamadryas Albers, 1850 : synonym of Drymaeus Albers, 1850
 * Hannarabdotus Emerson & Jacobson, 1964 : synonym of Rabdotus Albers, 1850
 * Kionoptyx Haas, 1966 : synonym of Bostryx Troschel, 1847 (junior synonymy)
 * Kuschelenia Hylton Scott, 1951 : synonym of Bocourtia (Kuschelenia) Hylton Scott, 1951 represented as Bocourtia Rochebrune, 1882
 * Leptodrymaeus Pilsbry, 1946 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
 * Leptomerus Albers, 1850 : synonym of Bulimulus Leach, 1814
 * Leptomormus Weyrauch, 1958 : synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
 * Liostracus [sic]: synonym of Leiostracus Albers, 1850 (misspelling of original genus, Leiostracus Albers, 1850 )
 * Loboa Ihering, 1917 : synonym of Bulimulus Leach, 1814
 * Mormus E. von Martens, 1860 : synonym of Drymaeus Albers, 1850
 * Multifasciatus Weyrauch, 1958 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
 * Naesiotellus Weyrauch, 1967 : synonym of Bostryx Troschel, 1847 (junior synonym)
 * Navicula Spix, 1827 : synonym of Cochlorina Jan, 1830
 * Nesiotes E. von Martens, 1860 : synonym of Naesiotus Albers, 1850
 * Obstrussus Parodiz, 1946 : synonym of Naesiotus Albers, 1850
 * Olinodia Dall, 1920 : synonym of Naesiotus (Reclasta) Dall, 1920 represented as Naesiotus Albers, 1850
 * Orphaicus Schaufuss, 1869 : synonym of Thaumastus E. von Martens, 1860
 * Orphnus Albers, 185 0: synonym of Thaumastus E. von Martens, 1860
 * Orthotomium Crosse & P. Fischer, 1875 : synonym of Rabdotus Albers, 1850
 * Pachyotus H. Beck, 1837 : synonym of Auris Spix, 1827
 * Pachytholus Strebel, 1909 : synonym of Thaumastus E. von Martens, 1860
 * Pampasinus Weyrauch, 1958 : synonym of Bostryx (Platybostryx) Pilsbry, 1896 : synonym ofBostryx Troschel, 1847
 * Peronaeus Albers, 1850 : synonym of Bostryx Troschel, 1847
 * Phenacotaxus Dall, 1912 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
 * Plecochilus Agassiz, 1846 : synonym of Plekocheilus Guilding, 1827 (unjustified emendation)
 * Pseudoperonaeus Weyrauch, 1958 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
 * Pseudorhodea Dall, 1895 : synonym of Rabdotus (Plicolumna) J.G. Cooper, 1895 represented as Rabdotus Albers, 1850
 * Pyrgus Albers, 1850 : synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
 * Rhabdotus [sic]: synonym of Rabdotus Albers, 1850 (incorrect subsequent spelling)
 * Rhaphiellus L. Pfeiffer, 1856 : synonym of Naesiotus Albers, 1850
 * Rimatula Parodiz, 1946 : synonym of Protoglyptus Pilsbry, 1897
 * Saeronia Dall, 1920 : synonym of Naesiotus Albers, 1850
 * Semiclausaria L. Pfeiffer, 1856 : synonym of Drymaeus Albers, 1850 (junior synonym)
 * Siphalomphix Rafinesque, 1833 : synonym of Bulimulus Leach, 1814
 * Sonorina Pilsbry, 1896 : synonym of Rabdotus (Leptobyrsus) Crosse & P. Fischer, 1875 represented as Rabdotus Albers, 1850
 * Spiroscutalus Pilsbry, 1932 : synonym of Scutalus Albers, 1850
 * Stenostoma Spix, 1827 : synonym of Anctus E. von Martens, 1860
 * Tatutor Jousseaume, 1887 : synonym of Thaumastus E. von Martens, 1860
 * Teneritia J. Mabille, 1898 : synonym of Spartocentrum Dall, 1895
 * Tholus Strebel, 1909 : synonym of Thaumastus E. von Martens, 1860
 * Xenothauma Fulton 1896: synonym of Scutalus Albers, 1850
 * Zaplagius Pilsbry, 1896 : synonym of Cochlorina Jan, 1830