Chipping sparrow

The chipping sparrow (Spizella passerina) is a species of New World sparrow, a passerine bird in the family Passerellidae. It is widespread, fairly tame, and common across most of its North American range.

There are two subspecies, the eastern chipping sparrow and the western chipping sparrow. This bird is a partial migrant with northerly populations flying southwards in the fall to overwinter in Mexico and the southern United States, and flying northward again in spring.

It molts twice a year. In its breeding plumage it has orangish-rust upper parts, gray head and underparts and a distinctive reddish cap. In non-breeding plumage, the cap is brown and the facial markings are less distinct. The song is a trill and the bird has a piercing flight call that can be heard while it is migrating at night.

In the winter, chipping sparrows are gregarious and form flocks, sometimes associating with other bird species. They mostly forage on the ground for seeds and other food items, as well as clambering on plants and trees, feeding on buds and small arthropods. In the west of their range they breed mainly in coniferous forests, but in the east, they choose woodland, farmland, parks and gardens. Breeding starts in late April and May and the nest is often built in a tree.

Description
Throughout the year, adults are gray below and an orangish-rust color above. Adults in alternate (breeding) plumage have a reddish cap, a nearly white supercilium, and a black trans-ocular line (running through the eye). Adults in basic (nonbreeding) plumage are less prominently marked, with a brownish cap, a dusky eyebrow, and a dark eye-line.

Juvenile chipping sparrows are prominently streaked below. Like non-breeding adults, they show a dark eye-line, extending both in front of and behind the eye. The brownish cap and dusky eyebrow are variable but generally obscure in juveniles.

Measurements:


 * Length: 4.7 - 5.9 in
 * Weight: 0.4 - 0.6 oz
 * Wingspan: 8.3 in

Vocalizations
The song is a trill that varies considerably among birds within any particular region. Two broad classes of variation in the song of the chipping sparrow are the fast trill and the slow trill. Individual elements in the fast trill are run together about twice as fast as in the slow trill; the fast trill sounds like a buzz or like someone snoring, whereas the slow trill sounds like rapid finger-tapping. Individual elements in the trill are very similar to a high pitch chi chi chi call.

The flight call of the chipping sparrow is heard year-round. Its flight call is piercing and pure-tone, lasting about 50 milliseconds. It starts out around 9 kHz, then falls to 7 kHz, then rises again to 9 kHz. The flight call may be transliterated as seen? Chipping sparrows migrate by night, and their flight calls are a characteristic sound of the night sky in spring and fall in the United States. In the southern Rockies and eastern Great Plains, the chipping sparrow appears to be the most common nocturnal migrant, judged by the number of flight calls detected per hour. On typical nights in August in this region, chipping sparrows may be heard at a rate of 15 flight calls per hour. On better-than-average nights, chipping sparrows occur at a rate of 60 flight calls per hour, and on exceptional nights chipping sparrows' flight calls are heard more than 200 times per hour.

Taxonomy
Chipping sparrows vary across their extensive North American range. There is minor geographic variation in appearance, and there is significant geographic variation in behavior. Ornithologists often divide the chipping sparrow into two major groups: the eastern chipping sparrow and the western chipping sparrow. However, there is additional plumage and behavioral variation within the western group.

At least two subspecies of chipping sparrows occur in western North America. The widespread Spizella passerina arizonae is associated with mountains and arid habitats of the western interior. A Pacific slope population constitutes subspecies S. p. stridula. Although these two races are both western, and are often lumped together as the western chipping sparrow, they do not necessarily form a single entity that stands apart from the eastern chipping sparrow (S. p. passerina).

Breeding


The male chipping sparrows start arriving at the breeding grounds from March (in more southern areas, such as Texas) to mid-May (in southern Alberta and northern Ontario). The female arrives one to two weeks later, and the male starts singing soon after to find and court a mate. After pair formation, nesting begins (within about two weeks of the female's arrival). Overall, the breeding season is from March till about August.

The chipping sparrow breeds in grassy, open woodland clearings and shrubby grass fields. The nest is normally above ground but below 6 m in height, and about 1 m on average, in a tree (usually a conifer, especially those that are young, short, and thick) or bush. The nest itself is constructed by the female in about four days. It consists of a loose platform of grass and rootlets and open inner cup of plant fiber and animal hair.

The chipping sparrow lays a clutch of two to seven pale blue to white eggs with black, brown, or purple markings. They are about 17 by, and incubated by the female for 10 to 15 days. The chipping sparrow is often brood parasitized by brown-headed cowbirds, usually resulting in the nest being abandoned.

Feeding
The chipping sparrow feeds on seeds year-round, although insects form most of the diet in the breeding season. Spiders are sometimes taken. Taraxacum officinale seeds are important during spring, and seeds from Fallopia convolvulus, Melilotus spp., Stellaria media, Chenopodium album, Avena spp., and others.

Throughout the year, chipping sparrows forage on the ground in covered areas, often near the edges of fields.

Monograph

 * Middleton, A. L. 1998. Chipping Sparrow (Spizella passerina), in: A. Poole and F. Gill, eds. The Birds of North America No. 334. The Birds of North America, Inc., Philadelphia, PA.

Thesis

 * Liu W-C. Ph.D. (2001). Development, variation, and use of songs by chipping sparrows. University of Massachusetts Amherst, United States, Massachusetts.
 * Perez DE. M.A. (1982). Parental Foraging in Chipping Sparrows (Spizella passerina). Western Michigan University, United States, Michigan.
 * Tate DJR. Ph.D. (1973). Habitat usage by the Chipping Sparrow (Spizella passerina) in northern lower Michigan. The University of Nebraska – Lincoln, United States, Nebraska.

Articles

 * Albrecht DJ & Oring LW. (1995). Song in chipping sparrows, Spizella passerina: Structure and function. Animal Behaviour. vol 50, pp. 1233–1241.
 * Anderson SH & Van Hook RIJ. (1973). Uptake and Biological Turnover of Cadmium-109 in Chipping Sparrows Spizella-Passerina. Environmental Physiology & Biochemistry. vol 3, no 5. pp. 243–247.
 * Braestrup FW. (1968). Evolution of Vertebrates Parus-Ater Parus-Cristatus Parus-Montanus Nemeritis-Canescens Phylloscopus-Sibilatrix Delichon-Urbica Spizella-Passerina Peromyscus-Maniculatus Turdus-Merula Turdus-Viscivorus Acrocephalus-Palustris. Zoologischer Anzeiger. vol 181, no 1/2. pp. 1–22.
 * Catherine PO & Joseph CO. (2001). Effects of Brown-headed Cowbirds on the nesting success of Chipping Sparrows in southwest Colorado. The Condor. vol 103, no 1. p. 127.
 * Dawson WR, Carey C, Adkisson CS & Ohmart RD. (1979). Responses of Brewers Sparrows Spizella-Breweri and Chipping Sparrows Spizella-Passerina to Water Restriction. Physiological Zoology. vol 52, no 4. pp. 529–541.
 * Earley CG. (1991). Brown-headed Cowbird, Molothrus ater, seen removing a Chipping Sparrow, Spizella passerina, egg. Canadian Field-Naturalist. vol 105, no 2. pp. 281–282.
 * Fillmore ER & Titman RD. (1977). CHIPPING SPARROW HANGED. Canadian Field-Naturalist. vol 91, no 1. pp. 69–69.
 * Foster J & Tozer R. (2001). Chipping sparrow feeds young of Eastern Kingbird. Ontario Birds. vol 19, no 2. pp. 79–83.
 * Liu W-C & Kroodsma DE. (1999). Song development by chipping sparrows and field sparrows. Animal Behaviour. vol 57, p. 1275.
 * Liu W-C & Kroodsma DE. (2007). Dawn and daytime singing behavior of chipping sparrows (Spizella passerina). Auk. vol 124, no 1. pp. 44–52.
 * Middleton ALA & Prescott DRC. (1989). POLYGYNY, EXTRA-PAIR COPULATIONS, AND NEST HELPERS IN THE CHIPPING SPARROW, SPIZELLA-PASSERINA. Canadian Field-Naturalist. vol 103, no 1. pp. 61–64.
 * Moldenha.Rr & Taylor PG. (1973). ENERGY-INTAKE BY HYDROPENIC CHIPPING SPARROWS (SPIZELLA-PASSERINA-PASSERINA) MAINTAINED ON DIFFERENT DIETS. Condor. vol 75, no 4. pp. 439–445.
 * Pulliam HR. (1980). Do Chipping Sparrows Spizella-Passerina-Arizonae Forage Optimally. Ardea. vol 68, no 1-4. pp. 75–82.
 * Reynolds JD & Knapton RW. (1984). Nest-Site Selection and Breeding Biology of the Chipping Sparrow Spizella-Passerina. Wilson Bulletin. vol 96, no 3. pp. 488–493.
 * Scott DM. (1988). HOUSE SPARROW AND CHIPPING SPARROW FEED THE SAME FLEDGLING BROWN-HEADED COWBIRD. Wilson Bulletin. vol 100, no 2. pp. 323–324.
 * Simmons GA & Sloan NF. (1974). Consumption of Jack-Pine Budworm Choristoneura-Pinus by the Eastern Chipping Sparrow Spizella-Passerina. Canadian Journal of Zoology. vol 52, no 7. pp. 817–821.
 * Sloan NF & Simmons GA. (1973). Foraging Behavior of the Chipping Sparrow in Response to High Populations of Jack Pine Budworm. American Midland Naturalist. vol 90, no 1. pp. 210–215.
 * Stewart PA. (1968). Bird Migration through an Abandoned Farmstead Richmondena-Cardinalis Behavior Dendroica-Palmarum Guiraca-Caerulea Spizella-Passerina. Chat. vol 32, no 4.
 * Swanson HM, Kinney B & Cruz A. (2004). Breeding biology of the Chipping Sparrow in ponderosa pine forests of the Colorado Front Range. Wilson Bulletin. vol 116, no 3. pp. 246–251.
 * Wan-Chun L. (2004). The effect of neighbours and females on dawn and daytime singing behaviours by male chipping sparrows. Animal Behaviour. vol 68, p. 39.
 * Wan-Chun L & Donald EK. (2006). SONG LEARNING BY CHIPPING SPARROWS: WHEN, WHERE, AND FROM WHOM. The Condor. vol 108, no 3. p. 509.
 * Zink RM & Dittmann DL. (1993). Population structure and gene flow in the chipping sparrow and a hypothesis for evolution in the genus Spizella. The Wilson Bulletin. vol 105, no 3. pp. 399–413.

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