Cochlostoma

Cochlostoma is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Cochlostomatidae.

Distribution
Calcareous terrains of South Europe and North Africa.

Description and Anatomy
Mantle caves serve as pulmones. Operculum corneous and multispiral. Penis is oriented above the right tentacle. Sexual dimorphism is often recognisable in external morphology.

Ecology
All species are confined to limestone habitats, feeding on dead plants and microflora. Females lays large eggs, hatching after three to six weeks.

Species
The genus Cochlostoma contains at least 140 species and subspecies, including:

subgenus Auritus
 * Cochlostoma achaicum (O. Boettger, 1885)
 * Cochlostoma alleryanum (Paulucci, 1879)
 * Cochlostoma auritum (Rossmässler, 1837)
 * Cochlostoma fuchsi Feher, 2004
 * Cochlostoma georgi (A.J. Wagner, 1906)
 * Cochlostoma hellenicum (Saint-Simon, 1869)
 * Cochlostoma hoyeri (Polinski, 1922)
 * Cochlostoma paladilhianum (Saint-Simon, 1869)
 * Cochlostoma parnonis Schütt, 1981
 * Cochlostoma pinteri Feher, 2004
 * Cochlostoma roseoli (A.J. Wagner, 1901)
 * Cochlostoma tessellatum (Rossmässler, 1837)
 * Cochlostoma westerlundi (Paulucci, 1879)

subgenus Cochlostoma
 * Cochlostoma septemspirale (Razoumowsky, 1789)

subgenus Turritus subgenus Dalfreddia
 * Cochlostoma acutum (Caziot, 1908)
 * Cochlostoma adamii (Paulucci, 1879)
 * Cochlostoma affine (Benoit, 1882)
 * Cochlostoma braueri (A.J. Wagner, 1897)
 * Cochlostoma crosseanum (Paulucci, 1879)
 * Cochlostoma elegans (Clessin, 1879)
 * Cochlostoma euboicum (Westerlund, 1885)
 * Cochlostoma gracile (L. Pfeiffer, 1849)
 * Cochlostoma kleciaki (Braun, 1887)
 * Cochlostoma macei (Bourguignat, 1869)
 * Cochlostoma montanum (Issel, 1866)
 * Cochlostoma mostarensis (A.J. Wagner, 1906)
 * Cochlostoma nanum (Westerlund, 1879)
 * Cochlostoma pageti Klemm, 1962
 * Cochlostoma patulum (Draparnaud, 1801)
 * Cochlostoma sardoum (Westerlund, 1890)
 * Cochlostoma simrothi (Caziot, 1908)
 * Cochlostoma stossichi (Hirc, 1881)
 * Cochlostoma sturanii (A.J. Wagner, 1897)


 * Cochlostoma porroi (Strobel, 1850)


 * Cochlostoma subalpinum (Pini, 1884)

subgenus Clessiniella


 * Cochlostoma anomphale Boeckel, 1939
 * Cochlostoma stelucarum Zallot et al., 2021
 * Cochlostoma tergestinum (Westerlund, 1878)
 * Cochlostoma villae (Strobel, 1851)
 * Cochlostoma waldemari (A.J. Wagner, 1897)

subgenus Eupomatias


 * Cochlostoma henricae (Strobel, 1851)
 * Cochlostoma philippianum (Gredler, 1853)

subgenus Wagneriola


 * Cochlostoma scalarinum (A. & J.B. Villa, 1841)

subgenus Lovcenia


 * Cochlostoma dalmatinum (L. Pfeiffer, 1863)
 * Cochlostoma erika (A.J. Wagner, 1906)
 * Cochlostoma jakschae Zallot, Fehér, Bamberger, Gittenberger, 2015
 * Cochlostoma lanatum Zallot, Fehér, Bamberger, Gittenberger, 2015
 * Cochlostoma tropojanum Zallot, Fehér, Bamberger, Gittenberger, 2015

unknown subgenus


 * Cochlostoma cinerascens (Rossmässler, 1837)
 * Cochlostoma cretense (Maltzan, 1887)

Synonyms of Cochlostoma

 * Auritus Westerlund, 1883
 * Auritus (Auritus) Westerlund, 1883
 * Auritus (Holcopoma) Kobelt & Möllendorff, 1899
 * Auritus (Pleuropoma) A. J. Wagner, 1897
 * Auritus (Titanopoma) A. J. Wagner, 1897
 * Cochlostoma (Auritus) Westerlund, 1883· accepted, alternate representation
 * Cochlostoma (Cinerascens) A. J. Wagner, 1897· accepted, alternate representation
 * Cochlostoma (Clessiniella) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015 (junior objective synonym)
 * Cochlostoma (Cochlostoma) Jan, 1830· accepted, alternate representation
 * Cochlostoma (Dalfreddia) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015 · accepted, alternate representation
 * Cochlostoma (Eupomatias) A. J. Wagner, 1897· accepted, alternate representation
 * Cochlostoma (Holcopoma) Kobelt & Möllendorff, 1899
 * Cochlostoma (Lovcenia) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015 · accepted, alternate representation
 * Cochlostoma (Neglecta) A. J. Wagner, 1897· accepted, alternate representation
 * Cochlostoma (Scalarina) A. J. Wagner, 1897· accepted, alternate representation
 * Cochlostoma (Titanopoma) A. J. Wagner, 1897 (junior synonym)
 * Cochlostoma (Turritus) Westerlund, 1883 · accepted, alternate representation
 * Cyclostoma (Cochlostoma) Jan, 1830 (original rank)
 * Diversicolor A. J. Wagner, 1905 (a junior synonym)
 * Hartmannia Newton, 1891
 * Holcopoma Kobelt & Möllendorff, 1899
 * Patuliana Caziot, 1908
 * Personatus Westerlund, 1883
 * Pleuropoma A. J. Wagner, 1897
 * Pleuropomatia Tomlin, 1929
 * Pomatias (Atlantica) A. J. Wagner, 1897 (invalid: junior homonym of Atlantica Ancey, 1887)
 * Pomatias (Auritus) Westerlund, 1883
 * Pomatias (Cinerascens) A. J. Wagner, 1897
 * Pomatias (Cisalpina) A. J. Wagner, 1897
 * Pomatias (Difficilis) A. J. Wagner, 1897 (invalid:junior objective synonym of Hartmannia)
 * Pomatias (Eupomatias) A. J. Wagner, 1897
 * Pomatias (Maculatus) Westerlund, 1883
 * Pomatias (Nana) A. J. Wagner, 1897 (invalid: junior homonym of Nana Schumacher, 1817)
 * Pomatias (Neglecta) A. J. Wagner, 1897
 * Pomatias (Personatus) Westerlund, 1883
 * Pomatias (Philippiana) A. J. Wagner, 1897
 * Pomatias (Pleuropoma) A. J. Wagner, 1897 ( invalid: junior homonym of Pleuropoma Möllendorff, 1893; * * Holcopoma and Pleuropomatia are replacement names)
 * Pomatias (Pomatiella) Clessin, 1889
 * Pomatias (Sardoa) A. J. Wagner, 1897
 * Pomatias (Scalarina) A. J. Wagner, 1897
 * Pomatias (Septemspiraliana) Caziot, 1910
 * Pomatias (Septemspiralis) A. J. Wagner, 1897
 * Pomatias (Strobelia) Clessin, 1889 (invalid: junior homonym of Strobelia Rondani, 1868 [Diptera])
 * Pomatias (Tesselata) A. J. Wagner, 1897
 * Pomatias (Titanopoma) A. J. Wagner, 1897 (junior objective synonym of Auritus)
 * Pomatias (Turritus) Westerlund, 1883
 * Stereopoma A. J. Wagner, 1897 (junior synonym)