Columella (genus)

Columella is a genus of very small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Truncatellinidae, the whorl snails and allies.

Taxonomy
Columella is commonly classified within the Vertiginidae. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the Chondrinidae or form a pupilloid family of its own. A systematic revision is pending.

Species
Species within the genus Columella include:
 * Columella acicularis Almuhambetova, 1979
 * Columella aspera Waldén, 1966
 * Columella columella (G. v. Martens, 1830)
 * Columella edentula (Draparnaud, 1805) - toothless column
 * Columella hartmutnordsiecki Schlickum & Geissert, 1980
 * Columella hasta (Hanna, 1911)
 * Columella intermedia Schileyko & Almuhambetova, 1984
 * Columella microspora (R.T. Lowe, 1852)
 * Columella nymphaepratensis Hlaváč & Pokryszko, 2009
 * Columella polvonense (Pilsbry, 1894)
 * Columella simplex (Gould, 1840)
 * Columella talgarica Schileyko & Rymzhanov, 2010
 * Columella wyciski Schlickum & Strauch, 1972
 * Species brought into synonymy:
 * Columella inornata (Michaud, 1831) : synonym of Columella edentula (Draparnaud, 1805) (junior synonym)
 * Columella ninagongonis Pilsbry, 1935 : synonym of Truncatellina ninagongonis (Pilsbry, 1935) (original combination)
 * Columella tridentata A. B. Leonard, 1946 : synonym of Gastrocopta ruidosensis (Cockerell, 1899)