Cychrus

Cychrus is a large genus of rare snail-eating beetles in the family Carabidae. There are at least 160 described species in Cychrus. They are found throughout the world, although more than 80% of the species occur in China.

Cychrus species
These 160 species belong to the genus Cychrus.


 * Cychrus aeneus Fischer von Waldheim, 1824 (Palearctic)
 * Cychrus altitudinum Deuve, 1998 (China)
 * Cychrus anatolicus Motschulsky, 1866 (Turkey)
 * Cychrus angulicollis Sella, 1874 (France and Italy)
 * Cychrus angustatus Hoppe & Hornschuch, 1825 (worldwide)
 * Cychrus angustior Kleinfeld, 2000 (China)
 * Cychrus angustitarsalis Deuve, 1991 (China)
 * Cychrus anomalognathus Imura & Cavazzuti, 2018 (China)
 * Cychrus attenuatus (Fabricius, 1792) (worldwide)
 * Cychrus auvrayorum Deuve & Mourzine, 1997 (China)
 * Cychrus barkamensis Deuve, 1991 (China)
 * Cychrus baxiensis Deuve, 1997 (China)
 * Cychrus becvari Deuve, 2018 (China)
 * Cychrus benesi Deuve, 1996 (China)
 * Cychrus bisetosus Deuve, 1995 (China)
 * Cychrus bispinosus Deuve, 1989 (China)
 * Cychrus boulbeni Deuve, 1997 (China)
 * Cychrus bruggei Deuve, 1991 (China)
 * Cychrus busatoi Cavazzuti, 2009 (China)
 * Cychrus businskyianus Imura, 2000 (China)
 * Cychrus businskyorum Imura, 2000 (China)
 * Cychrus caraboides (Linnaeus, 1758) (snail hunter)  (worldwide)
 * Cychrus cavazzutii Deuve, 2002 (China)
 * Cychrus cerberus Cavazzuti, 2007 (China)
 * Cychrus chareti Deuve, 1993 (China)
 * Cychrus colasi Deuve, 1994 (China)
 * Cychrus coptolabroides Imura & Cavazzuti, 2017 (China)
 * Cychrus cordicollis Chaudoir, 1835 (Europe)
 * Cychrus cordithorax Deuve, 2007 (China)
 * Cychrus cyaneomelas Deuve, 2010
 * Cychrus cylindricollis Pini, 1871 (Italy)
 * Cychrus dacatrai Deuve, 1992 (China)
 * Cychrus daochengicus Deuve, 1989 (China)
 * Cychrus davidis Fairmaire, 1886 (China)
 * Cychrus deuveianus Cavazzuti, 1998 (China)
 * Cychrus dolichognathus Deuve, 1990 (China)
 * Cychrus dufouri Chaudoir, 1869 (France and Spain)
 * Cychrus elongaticeps Deuve, 1992 (China)
 * Cychrus elongatulus Deuve, 2001 (China)
 * Cychrus evae Häckel & Sehnal, 2007 (China)
 * Cychrus forceps Imura & Cavazzuti, 2017 (China)
 * Cychrus fugongensis Imura & Cavazzuti, 2018 (China)
 * Cychrus furumii Deuve, 1990 (China)
 * Cychrus gorodinskiellus Deuve, 2016 (China)
 * Cychrus gorodinskiorum Deuve, 2013 (China)
 * Cychrus grajus K. & J.Daniel, 1898 (Italy)
 * Cychrus grumulifer Deuve, 1994 (China)
 * Cychrus haesitans Cavazzuti, 2007 (China)
 * Cychrus hampei Gestro, 1875 (Bosnia-Herzegovina and Croatia)
 * Cychrus heishuiensis Deuve, 2004 (China)
 * Cychrus hemphillii G. Horn, 1879 (North America)
 * Cychrus horribilis Deuve, 2016 (China)
 * Cychrus huyaensis Imura & Cavazzuti, 2018 (China)
 * Cychrus inexpectatior Deuve, 1991 (China)
 * Cychrus inframontes Cavazzuti, 2007 (China)
 * Cychrus inops Deuve, 1997 (China)
 * Cychrus italicus Bonelli, 1810 (Europe)
 * Cychrus janatai Deuve, 2001 (China)
 * Cychrus jiaolieshanus Imura & Cavazzuti, 2019 (China)
 * Cychrus jinchuanensis Deuve & Tian, 2007 (China)
 * Cychrus jinpingshanus Imura & Cavazzuti, 2017 (China)
 * Cychrus jirouxi Deuve, 2006 (China)
 * Cychrus jiulongensis Deuve, 1994 (China)
 * Cychrus kabakianus Imura & Cavazzuti, 2017 (China)
 * Cychrus kalabi Deuve, 1991 (China)
 * Cychrus kalabianus Deuve, 2019 (China)
 * Cychrus kangensis Deuve & Tian, 2013 (China)
 * Cychrus kaznakovi Semenov & Znojko, 1934 (China)
 * Cychrus keithi Deuve, 1998 (China)
 * Cychrus koiwayai Deuve & Imura, 1993 (China)
 * Cychrus korotyaevi Deuve, 1991 (China)
 * Cychrus kozlovi Semenov & Znojko, 1934 (China)
 * Cychrus kralianus Deuve, 1996 (China)
 * Cychrus kryzhanovskii Deuve, 2000 (China)
 * Cychrus kubani Deuve, 1992 (China)
 * Cychrus kvetoslavae Deuve, 2006 (China)
 * Cychrus lajinensis Deuve & Tian, 2002 (China)
 * Cychrus lanpingensis Deuve, 1997 (China)
 * Cychrus lecordieri Deuve, 1990 (China)
 * Cychrus lianghensis Cavazzuti, 2008 (China)
 * Cychrus liei Kleinfeld, 2003 (China)
 * Cychrus lilianae Cavazzuti, 1997 (China)
 * Cychrus liuyei Deuve & Tian, 2013 (China)
 * Cychrus loccai Cavazzuti, 1997 (China)
 * Cychrus luctifer Deuve, 1991 (China)
 * Cychrus ludmilae Imura, 1999 (China)
 * Cychrus luhuo Deuve, 1994 (China)
 * Cychrus luojiensis Cavazzuti, 1998 (China)
 * Cychrus luolaluokuanus Imura & Cavazzuti, 2019 (China)
 * Cychrus maoxianicus Deuve & Mourzine, 2000 (China)
 * Cychrus marcilhaci Deuve, 1992 (China)
 * Cychrus minjiangicus Deuve, 2009 (China)
 * Cychrus minshanicola Deuve, 1987 (China)
 * Cychrus miroslavi Deuve, 2006 (China)
 * Cychrus moerkuaicus Deuve & Tian, 2007 (China)
 * Cychrus morawitzi Géhin, 1885 (Asia)
 * Cychrus morvani Deuve, 1998 (China)
 * Cychrus motianling Imura & Cavazzuti, 2018 (China)
 * Cychrus mugecuo Deuve, 1994 (China)
 * Cychrus muliensis Deuve, 1992 (China)
 * Cychrus naviauxi Deuve & Mourzine, 1998 (China)
 * Cychrus nixiensis Imura & Cavazzuti, 2018 (China)
 * Cychrus ombrophilus Deuve, 1990 (China)
 * Cychrus pangi Deuve & Tian, 2004 (China)
 * Cychrus paraxiei Cavazzuti, 2009 (China)
 * Cychrus pentaploidus Deuve, 2010 (China)
 * Cychrus pratti Breuning, 1946 (China)
 * Cychrus procerus Cavazzuti, 1998 (China)
 * Cychrus prosciai Cavazzuti, 2010 (China)
 * Cychrus pseudokoiwayai Deuve, 1998 (China)
 * Cychrus puchneri Kleinfeld, 2016 (China)
 * Cychrus puetzi Kleinfeld & Korell&Wrase, 1996 (China)
 * Cychrus quadrisetifer Imura, 1998 (China)
 * Cychrus remondi Deuve, 1999 (China)
 * Cychrus reni Deuve & Tian, 2013 (China)
 * Cychrus reuteri Kleinfeld, 2018 (China)
 * Cychrus riwaensis Deuve, 2006 (China)
 * Cychrus rosti Roeschke, 1907 (Georgia and Russia)
 * Cychrus rostislavi Deuve, 2009 (China)
 * Cychrus rugicollis K. & J.Daniel, 1898 (Bosnia-Herzegovina)
 * Cychrus rugocephalus Deuve, 2014 (China)
 * Cychrus sabdensis Deuve, 1994 (China)
 * Cychrus sars Imura & Häckel, 2003 (China)
 * Cychrus schmidtii Chaudoir, 1837 (Europe)
 * Cychrus schneideri Imura, 1997 (China)
 * Cychrus sehnali Häckel, 2007 (China)
 * Cychrus sellemi Deuve, 2002 (China)
 * Cychrus semelai Deuve, 1997 (China)
 * Cychrus semigranosus Palliardi, 1825 (worldwide)
 * Cychrus seriatus Roeschke, 1907 (China)
 * Cychrus shangrilanus Imura & Cavazzuti, 2017 (China)
 * Cychrus shankoucola Deuve, 1994 (China)
 * Cychrus shanxiensis Deuve, 2005 (China)
 * Cychrus signatus Faldermann, 1836 (Georgia and Turkey)
 * Cychrus sinicus Deuve, 1989 (China)
 * Cychrus sola Cavazzuti, 1997 (China)
 * Cychrus songpanensis Deuve, 1991 (China)
 * Cychrus spinicollis L.Dufour, 1857 (Spain)
 * Cychrus starcki Reitter, 1888 (Georgia and Russia)
 * Cychrus stoetzneri Roeschke, 1923 (China)
 * Cychrus szetshuanus Breuning, 1931 (China)
 * Cychrus thibetanus Fairmaire, 1893 (China)
 * Cychrus tubaemons Imura & Cavazzuti, 2017 (China)
 * Cychrus tuberculatus T. Harris, 1839 (North America)
 * Cychrus turnai Deuve, 1994 (China)
 * Cychrus uenoi Imura, 1995 (China)
 * Cychrus vignai Cavazzuti, 2011 (China)
 * Cychrus wuyipeng Deuve, 1992 (China)
 * Cychrus xiei Deuve, 1989 (China)
 * Cychrus yadingensis Deuve, 2006 (China)
 * Cychrus yajiangensis Deuve & Mourzine, 1998 (China)
 * Cychrus yi Cavazzuti, 2001 (China)
 * Cychrus yulongxuicus Deuve, 1991 (China)
 * Cychrus yunnanus Fairmaire, 1887 (China)
 * Cychrus zhimahensis Imura & Cavazzuti, 2019 (China)
 * Cychrus zhoui Imura & Su&Osawa, 1998 (China)
 * Cychrus zoigeicus Deuve, 1990 (China)
 * Cychrus zuopengensis Imura & Cavazzuti, 2019 (China)
 * † Cychrus minor G.Horn, 1876
 * † Cychrus wheatleyi G.Horn, 1876