Escucha Formation

The Escucha Formation is a geological formation in La Rioja and Teruel provinces of northeastern Spain whose strata date back to the late Aptian to middle Albian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.

The approximately 70 m thick formation underlies the Utrillas Formation and overlies Castrillo de la Reina, Benassal & Oliete Formations. The Escucha Formation comprises siltstones, mudstones, sandstones, coal, siltstones and amber, in which several fossil insects were found. The formation was deposited in a variety of continental to paralic (deltaic) environments.

Fossil content
The Escucha Formation has provided the following fossils, among others:

Dinosaurs

 * Other dinosaurs
 * Allosauroidea indet.
 * Iguanodontia indet.
 * Titanosauriformes indet.


 * Reptiles
 * Aragochersis lignitesta
 * Toremys cassiopeia
 * Trachydermochelys sp.
 * Testudines indet.
 * Anteophthalmosuchus escuchae
 * Goniopholididae indet.
 * Hulkepholis plotos


 * Fish
 * Chondrichthyes indet.
 * Osteichthyes indet.

Insects

 * Other arthropods


 * Alavia neli
 * Archaelagonops alavensis
 * Mesozygiella dunlopi
 * Alavesiaphis margaritae
 * Hispanocader lisae
 * Alavametra popovi
 * Iberovelia quisquilia
 * Glaesivelia pulcherrima
 * Gymnopollisthrips maior
 * G. minor
 * Manicapsocidus enigmaticus
 * Archaeatropos alavensis
 * Preempheria antiqua
 * Empheropsocus arilloi
 * E. margineglabrus
 * Morazatermes krishnai
 * Cantabritermes simplex
 * Autrigonoforceps iberica
 * Hispanelcana alavensis
 * H. arilloi
 * H. lopezvallei
 * Libanophron sugaar
 * Hippocoon basajauni
 * Burmaphron iratxoak
 * B. jentilak
 * B. sorginak
 * Tagsmiphron olentzero
 * Elasmophron mari
 * Ectenobythus iberiensis
 * Liztor pilosus
 * Cretepyris martini
 * Zophepyris alavaensis
 * Ampulicomorpha perialla
 * Microcostaphron parvus
 * Archephedrus stolamissus
 * Protorhyssalopsis perrichoti
 * Spathiopteryx alavarommopsis
 * Serphites lamiak
 * Aposerphites angustus
 * Archaeromma hispanicum
 * Galloromma alavaensis
 * Alavaromma orchamum
 * Tithonoscelio resinalis
 * Bruescelio platycephalus
 * Electroteleiopsis hebdomas
 * Alavascelio delvallei
 * Amissascelio temporarius
 * Perimoscelio tyrbastes
 * Perimoscelio confector
 * Proterosceliopsis masneri
 * Juxtascelio interitus
 * Iberopria perialla
 * Iberoevania roblesi
 * Cretevania alonsoi
 * Iberomaimetsha nihtmara
 * Iberomaimetsha rasnitsyni
 * Valaa delclosi
 * Megalava truncata
 * Eosyntexis parva
 * Cretasonoma corinformibus
 * Penarhytus tenebris
 * Prosolierius parvus
 * Cretakarenni hispanicus
 * Rhizophtoma longus
 * Darwinylus marcosi
 * Mediumiuga sinespinis
 * Archiaustroconops alavensis
 * Leptoconops (Leptoconops) zherikhini
 * Gerontodacus skalskii
 * Chimeromyia alava
 * Chimeromyina concilia
 * Euliphora grimaldii
 * Alavesia subiasi
 * Tethepomima holomma
 * Tethepomyia buruhandi
 * Lysistrata emerita
 * Hegalari minor
 * Hegalari antzinako
 * Alavamanota hispanica
 * Allocotocera xavieri
 * Cretohaplusia ortunoi
 * Eltxo cretaceus
 * Helius (Helius) alavensis
 * Helius (Helius) spiralis
 * Espanoderus barbarae
 * Alavaraphidia imperterrita
 * Baissoptera cretaceoelectra
 * Cretokatianna bucculenta
 * Sphyrotheciscus senectus
 * Archeallacma dolichopoda
 * Katiannasminthurus xenopygus
 * Pseudosminthurides stoechus
 * Burmisotoma spinulifera
 * Protoisotoma autrigoniensis
 * Proisotoma communis
 * Proleptochelia tenuissima
 * Electrotanais monolithus
 * Alavatanais carabe
 * Alavatanais margulisae
 * Eurotanais terminator
 * Spinomegops arcanus
 * Strieremaeus minguezae
 * Iberofoveopsis miguelesi
 * Hispanothrips utrillensis
 * Aragonitermes teruelensis
 * Aragonimantis aenigma
 * Mymaropsis turolensis
 * Serphites silban
 * Galloromma turolensis
 * Alavaromma orchamum
 * Cretevania rubusensis
 * Cretevania alcalai
 * Cretevania montoyai
 * Actenobius magneoculus
 * Arra legalovi
 * Leptoconops (Leptoconops) zherikhini
 * Atriculicoides sanjusti
 * Atriculicoides hispanicus
 * Gerontodacus skalskii
 * Microphorites utrillensis
 * Litoleptis fossilis
 * Burmazelmira grimaldii
 * Aragomantispa lacerata
 * Spinomegops aragonensis
 * Ametroproctus valeriae
 * Cretaceobodes martinezae
 * Trhypochthonius lopezvallei


 * Invertebrates
 * Ostracoda indet.
 * Bivalvia indet.
 * Gastropoda indet.


 * Flora
 * Pinophyta
 * Angiospermae indet.