Fossil flora of Kızılcahamam district

This fossil flora in Turkey stems from at least six Pliocene deposits in Güvem and Beşkonak villages, 80 km north of Ankara, 22 km north of Kızılcahamam and 125 km south of the Black Sea coast. They are 1000 m to 1500 m above sea level, extending 15 km north-south and 7 km east-west. Between six and seven million years ago, a fresh water lake existed there in a forested area with mostly broad leaved deciduous tree species, dominated by sequoia and oak.

The Turkish botany Professor Baki Kasaplıgil (1918-1992) from Çankırı who worked at the Biology Department, Mills College, Oakland, California from 1956 to 1992, made fossil collecting trips to the area around Kızılcahamam - Çerkeş road in 1964 and 1968. In the 1964 trip he collected fossil leaves of Glyptostrobus, Sequoia, Zelkova, Fagus, Carpinus, Ulmus, Quercus, Tilia and Acer. In the 1968 trip, he collected 400 plant parts such as leaves, seeds, fruits, twigs and branches. He also collected 50 fossils of frog and fish skeletons and larvae and insects. In 1969, Wayne Fry from University of Berkeley made plant fossil collections in Güvem. Daniel I. Axelrod also assisted in the research of this flora.

Pteridophyta

 * Salviniaceae


 * Salvinia sp.


 * Pteridaceae


 * Cryptogramma aff. crispo

Gymnosperms

 * Cupressaceae


 * †Glyptostrobus europaeus


 * †Sequoia langsdorfi


 * Pinaceae


 * Pinus canariensis


 * Pinus morrisonicola


 * Tsuga sp.


 * Ephedraceae


 * Ephedra sp.

Angiosperms, dicots

 * Cercidiphyllaceae


 * †Cercidiphyllum crenatum


 * Magnoliaceae


 * Magnolia sprengeri


 * Menispermaceae


 * Menispermum sp.


 * Lauraceae


 * Persea indica


 * Platanaceae


 * Platanus sp.


 * Juglandaceae


 * †Platycarya miocenica


 * Pterocarya fraxinifolia


 * Salicaceae


 * Salix sp.


 * Populus tremula


 * Myricaceae


 * Comptonia sp.


 * Myrica banksiaefolia


 * Fagaceae


 * Castanea sp.


 * Castanopsis sp.


 * Fagus sp.


 * †Quercus drymeja
 * †Quercus heidingeri
 * †Quercus kubinyi
 * †Quercus aff. lonchitis
 * Quercus petroela aff.
 * Quercus sclerophyllina
 * Quercus semecarpifolia
 * †Quercus seyfriedii (very similar to Quercus phellos from eastern and central United States)
 * †Quercus sosnoisky
 * Quercus hartwissiana
 * Quercus trojana aff.


 * Betulaceae


 * Alnus sp.


 * Betula luminifera aff.


 * Carpinus miocenica


 * Sapindaceae


 * †Acer angustilobum
 * †Acer tribolatum


 * Anacardiaceae


 * Astronium sp.


 * Ebenaceae


 * Diospyros aff. miokaki


 * Aquifoliaceae


 * Ilex gracilis (close to Ilex serrata)


 * Rosaceae
 * Sorbus aucuparia aff.


 * Fabaceae


 * Cercis sp.


 * †Sophora sp.


 * Malvaceae


 * Tilia platyphyllos aff,


 * Ulmaceae


 * Ulmus sp.


 * †Zelkova ungeri


 * Berberidaceae


 * Berberis chinensis aff.


 * Moraceae


 * Ficus sp.


 * Altingiaceae


 * †Liquidambar europaeum


 * Simaroubaceae


 * Ailanthus altissima aff.

Angiosperms, monocots

 * Smilaceae


 * Smilax aspera


 * Potamogetonaceae


 * Potamogeton sp.


 * Hydrocharitaceae


 * Egeria densa aff.


 * Najas sp.


 * Typhaceae


 * Typha sp.