Frullania



Frullania is the only genus of liverworts in family Frullaniaceae. It contains the following species:

A
 * Frullania aculeata Taylor, 1846
 * Frullania acutata Caspary, 1887
 * Frullania acutiloba Gerola, 1947
 * Frullania akiyamae Hattori, 1986
 * Frullania albertii Stephani, 1916
 * Frullania allanii Hodgson, 1949
 * Frullania allionii Stephani, 1910
 * Frullania alpina Stephani, 1911
 * Frullania alstonii Verdoorn, 1930
 * Frullania alstonii var. pfleidereri Hattori, 1972
 * Frullania alternans Nees In G., L. & N., 1845
 * Frullania amamiensis Kamimura, 1968
 * Frullania ambronnii Stephani, 1916
 * Frullania amplicrania Stephani, 1910
 * Frullania ampullifera Jack & Stephani In Stephani, 1894
 * Frullania anderssonii Ångström, 1873
 * Frullania angulata Mitten, 1863
 * Frullania angulata F. Serratoides Vanden Berghen, 1983
 * Frullania angulata var. laciniata Demaret & Vanden Berghen, 1950
 * Frullania angustistipa Stephani, 1908
 * Frullania anomala Hodgson, 1949
 * Frullania antaresensis Hattori, 1980
 * Frullania aoshimensis Horikawa, 1929
 * Frullania apertilobula Gerola, 1947
 * Frullania apicalis Mitten, 1879
 * Frullania apicalis var. camerunensis Vanden Berghen, 1976
 * Frullania apiculata (R., B. & N.) Nees In G., L. & N., 1845
 * Frullania apiculata var. goebelii Schiffner, 1893
 * Frullania apiculata var. guianensis Lindenberg & Gottsche, 1851
 * Frullania apiculata var. laxa Nees In G., L. & N., 1845
 * Frullania apollinarii Stephani, 1911
 * Frullania aposinensis Hattori & Lin, 1985
 * Frullania appalachiana Schuster, 1983
 * Frullania appendistipula Hattori, 1972
 * Frullania appendistipula var. spinifera Hattori, 1974
 * Frullania arecae (Sprengel) Gottsche, 1863
 * Frullania arecae var. spiniloba (Stephani) Yuzawa, 1991
 * Frullania armata Herzog & Clark In Clark, 1954
 * Frullania armatifolia Verdoorn, 1932
 * Frullania armitiana Stephani, 1911
 * Frullania armitiana var. inflexula Hattori, 1988
 * Frullania armitiana var. longe-Attenuata (Hattori) Hattori, 1980
 * Frullania arsenii Stephani, 1924
 * Frullania asagrayana Montagne, 1842
 * Frullania astrolabea Stephani, 1910
 * Frullania aterrima (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
 * Frullania aterrima var. lepida Hodgson, 1949
 * Frullania atrata (Swartz) Montagne, 1839
 * Frullania atrata var. flaccida (Weber) Nees In G., L. & N., 1845
 * Frullania atrata var. mexicana Nees In G., L. & N., 1845
 * Frullania atrosanguinea Taylor Ex Spruce, 1884
 * Frullania attenuata Stephani, 1911
 * Frullania auriculata Hattori, 1985
 * Frullania azorica Sim-Sim, Sergio, Mues & Kraut, 1995

B
 * Frullania baileyana Stephani, 1910
 * Frullania bakeri Stephani, 1916
 * Frullania baladina Gottsche Ex Stephani, 1894
 * Frullania baladina var. edentata Hattori, 1984
 * Frullania baltica Grolle, 1985
 * Frullania baumannii Hattori, 1977
 * Frullania beauverdii Stephani, 1916
 * Frullania bella Stephani, 1911
 * Frullania belmorensis Stephani In Stephani & Watts, 1914
 * Frullania benjaminiana Inoue In Hattori, 1975
 * Frullania bergmanii Hattori, 1974
 * Frullania berthoumieui Stephani, 1894
 * Frullania beyrichiana (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania bhutanensis Hattori, 1971
 * Frullania bicornistipula Spruce, 1884
 * Frullania blastopetala Hattori, 1984
 * Frullania blepharozia Spruce, 1884
 * Frullania bogotensis Stephani, 1910
 * Frullania bolanderi Austin, 1869
 * Frullania bonariensis Reiner, 1988
 * Frullania bonincola Hattori, 1978
 * Frullania borbonica Lindenberg In G., L. & N., 1845
 * Frullania boveana Massalongo, 1885
 * Frullania brachycarpa Spruce, 1889
 * Frullania brasiliensis Raddi, 1822
 * Frullania brasiliensis var. cylindrica (Gottsche In Lehmann) Spruce, 1884
 * Frullania brasiliensis var. cylindrica F. Lindigii (Gottsche) Spruce, 1884
 * Frullania breuteliana Gottsche In G., L. & N., 1845
 * Frullania brevicalycina Stephani, 1894
 * Frullania brittoniae Evans, 1897
 * Frullania brotheri Stephani, 1894
 * Frullania brunnea (Sprengel) Drège, 1843
 * Frullania buchtienii Herzog, 1942
 * Frullania bullata Stephani, 1910

C
 * Frullania caduca Hattori, 1980
 * Frullania caespitans Beauverd In Stephani, 1924
 * Frullania caffraria Stephani, 1894
 * Frullania caffraria F. Anomala Vanden Berghen, 1976
 * Frullania calcarata Ångström, 1873
 * Frullania caldensis Ångström, 1876
 * Frullania caledonica Gottsche Ex Stephani, 1894
 * Frullania californica (Austin Ex Underwood) Evans, 1897
 * Frullania campanulata Sande Lacoste, 1853
 * Frullania campanulata var. caduca Verdoorn, 1930
 * Frullania campanulata var. malesiaca (Verdoorn) Hattori, 1975
 * Frullania canaliculata Gottsche Ex Stephani, 1910
 * Frullania capensis Gottsche In G., L. & N., 1845
 * Frullania capillaris Stephani, 1911
 * Frullania carrii Kamimura & Hattori In Hattori & Kamimura, 1973
 * Frullania casparyi Grolle, 1985
 * Frullania cataractarum Stephani, 1911
 * Frullania caulisequa (Nees In Martius) Montagne, 1839
 * Frullania cavallii Gola, 1907
 * Frullania cesatiana De Notaris, 1865
 * Frullania changii Hattori & Gao, 1985
 * Frullania chenii Hattori & Lin, 1985
 * Frullania chevalieri (Schuster) Schuster, 1992
 * Frullania chiapasana Stephani, 1910
 * Frullania chilcootiensis Stephani, 1887
 * Frullania chilensis Stephani, 1894
 * Frullania chiovendae Gola, 1914
 * Frullania chodatii Beauverd In Stephani, 1924
 * Frullania ciliata Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania cinchonae Gottsche In G., L. & N., 1845
 * Frullania clandestina (Nees & Montagne) Nees In G., L. & N., 1845
 * Frullania clavata (Hooker & Taylor) Taylor In G., L. & N., 1845
 * Frullania claviloba Stephani, 1911
 * Frullania clemensiana Verdoorn, 1932
 * Frullania cobrensis Gottsche Ex Stephani, 1894
 * Frullania colliculosa von Konrat, Braggins, Hentschel & Heinrichs, 2010
 * Frullania compacta Gottsche Ex Stephani, 1911
 * Frullania complicata Stephani, 1911
 * Frullania confertiloba Stephani, 1910
 * Frullania consociata Stephani, 1910
 * Frullania contracta Stephani, 1911
 * Frullania controversa Beauverd In Stephani, 1924
 * Frullania convoluta Lindenberg & Hampe In Hampe, 1851
 * Frullania convoluta var. ampliata Herzog, 1953
 * Frullania cordaeana Lindenberg In G., L. & N., 1845
 * Frullania cordistipula (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
 * Frullania cordistipula var. (Beta) mutica Nees In G., L. & N., 1845
 * Frullania cordistipula var. (Gamma) regularis Nees In G., L. & N., 1845
 * Frullania cordistipula var. dentistipula Hattori, 1986
 * Frullania cornuta Stephani, 1911
 * Frullania crassitexta Stephani, 1910
 * Frullania crawfordii Stephani, 1894
 * Frullania crenulifolia Jack & Stephani, 1892
 * Frullania crinoidea Spruce Ex Stephani In Stephani, 1911
 * Frullania crispiloba Stephani, 1894
 * Frullania crispiplicata Yuzawa & Hattori Ex Konstantinova, Potemkin & Schijakov, 1992
 * Frullania cristata Hattori, 1981
 * Frullania cuencensis Taylor, 1846
 * Frullania cuneatistipula Stephani, 1924
 * Frullania cuneiloba Nees In G., L. & N., 1845
 * Frullania curviramea Stephani, 1911
 * Frullania curvirostris Colenso, 1889
 * Frullania curvistipula Stephani, 1911
 * Frullania curvistipula var. falcatidenta Hattori, 1982
 * Frullania curvistipula var. lamii Verdoorn, 1930
 * Frullania curvistipula var. latistipula Hattori, 1978
 * Frullania cuspidifolia Stephani, 1911
 * Frullania cuspiloba Stephani, 1910
 * Frullania cyparioides (Schwägrichen) Schwaegrichen In G., L. & N., 1845

D
 * Frullania darwinii Gradstein & Uribe In Uribe, 2004
 * Frullania davurica Hampe In G., L. & N., 1845
 * Frullania davurica F. Dorsoblastos (Hattori) Hattori & Lin, 1985
 * Frullania davurica F. Microphylla (Massalongo) Hattori & Lin, 1985
 * Frullania davurica var. chichibuensis (Kamimura) Hattori, 1976
 * Frullania davurica var. concava Chang In Gao & Chang, 1981
 * Frullania debilis Stephani Ex Hattori, 1974
 * Frullania decidua Spruce, 1884
 * Frullania deflexa Mitten, 1862
 * Frullania degelii Arnell, 1959
 * Frullania densifolia Subsp. Andamana Hattori Ex Singh In Nath & Asthana, 2001
 * Frullania densiloba Stephani Ex Evans, 1906
 * Frullania dentata Hattori, 1974
 * Frullania dentata var. secernens Hattori, 1988
 * Frullania dentifera Hattori & Streimann, 1985
 * Frullania dentiloba Hattori, 1975
 * Frullania deplanata Mitten In Hooker, 1855
 * Frullania deppii Gottsche Ex Lehmann, 1844
 * Frullania depressa Mitten, 1863
 * Frullania dilatata Subsp. (Dilatata 1982) (L.) Dumortier, 1835
 * Frullania dilatata Subsp. (Dilatata 1982) F. Fuscovirens Jorgensen, 1934
 * Frullania dilatata Subsp. (Dilatata 1982) var. anomala Corbiere, 1889
 * Frullania dilatata Subsp. (Dilatata 1982) var. macrotus Nees, 1838
 * Frullania dilatata Subsp. (Dilatata 1982) var. subtilissima Nees, 1838
 * Frullania dilatata Subsp. Asiatica Hattori, 1982
 * Frullania diptera (Lehmann & Lindenberg) Drège, 1843
 * Frullania dispar Nees In G., L. & N., 1845
 * Frullania diversitexta Stephani, 1897
 * Frullania donnellii Austin, 1879
 * Frullania dulimensis Uribe, 2006
 * Frullania durifolia Stephani, 1894
 * Frullania dusenii Stephani In Dusen, 1905
 * Frullania duthiana Stephani, 1910
 * Frullania duthiana var. laevis Hattori In Ohashi, 1975
 * Frullania duthiana var. szechuanensis Hattori & Gao, 1985

E
 * Frullania eboracensis (Subsp. Eboracensis 1992) Gottsche Ex Lehmann, 1844
 * Frullania eboracensis Subsp. Virginica (Gottsche In Lehmann) Schuster, 1992
 * Frullania echinantha Hattori, 1974
 * Frullania echinatella Hattori, 1988
 * Frullania ecklonii (Sprengel) Sprengel In G., L. & N., 1845
 * Frullania ecklonii F. Robustior Gottsche, 1863
 * Frullania ecklonii F. Tenerior Sprengel, 1847
 * Frullania ecklonii var. huitamalcensis Gottsche, 1863
 * Frullania ecklonii var. rufescens Sprengel, 1847
 * Frullania ecuadoriensis Stephani, 1911
 * Frullania elegans Lehmann, 1857
 * Frullania elephantum Hattori, 1977
 * Frullania engelii Hattori, 1983
 * Frullania epiphylla Subsp. Fijiensis Hattori, 1985
 * Frullania eplicata Stephani, 1911
 * Frullania ericoides (Nees In Martius) Montagne, 1839
 * Frullania ericoides F. Ericoides (Nees) Verdoorn Ex Piippo, 1990
 * Frullania ericoides var. planescens (Verdoorn) Hattori, 1984
 * Frullania erostrata Hattori, 1974
 * Frullania errans Verdoorn, 1930
 * Frullania errans var. angulistipula Hattori, 1972
 * Frullania esenbeckiana Beauverd Ex Stephani, 1924
 * Frullania evelynae Hattori & Thaithong, 1978
 * Frullania evoluta Mitten, 1861
 * Frullania expansa Stephani, 1897
 * Frullania eymae Hattori, 1975
 * Frullania eymae var. crispidentata Hattori & Streimann, 1985

F
 * Frullania falciloba Taylor Ex Lehmann, 1844
 * Frullania fallax Gottsche In G., L. & N., 1845
 * Frullania falsicornuta Hattori, 1986
 * Frullania falsisinuata Hattori & Piippo, 1986
 * Frullania falsisinuata F. Parvistylata (Hattori) Hattori & Piippo, 1986
 * Frullania falsisinuata var. crispidentata Hattori & Piippo, 1986
 * Frullania fauriana Stephani, 1894
 * Frullania fauriana F. Emarginata Kamimura, 1952
 * Frullania fengyangshanensis Zhu & So, 1997
 * Frullania ferdinandi-Muelleri Stephani, 1910
 * Frullania fertilis var. major Massalongo, 1885
 * Frullania flammea Taylor Ex Spruce, 1884
 * Frullania flexicaulis Spruce, 1884
 * Frullania flexuosa Hattori, 1983
 * Frullania fragilifolia (Taylor) Taylor In G., L. & N., 1845
 * Frullania franciscana Howe, 1894
 * Frullania fuegiana Stephani, 1910
 * Frullania fugax (Hooker & Taylor) Taylor In G., L. & N., 1845
 * Frullania fulfordiae Hattori, 1987
 * Frullania fusco-Virens Stephani, 1910
 * Frullania fusco-Virens var. gemmipara (Schuster & Hattori In Hattori) Hattori & Lin, 1985

G
 * Frullania gabonensis Vanden Berghen, 1976
 * Frullania gaoligongensis Bai & Gao, 1999
 * Frullania gaudichaudii (Nees & Montagne) Nees & Montagne In G., L. & N., 1845
 * Frullania gaudichaudii F. Hasseltii (Sande Lacoste) Verdoorn, 1930
 * Frullania gemmulosa Hattori & Thaithong In Hattori, Thaithong & Kitagawa, 1977
 * Frullania gibbosa (Nees) Nees In Montagne, 1840
 * Frullania gibbosa var. major Gottsche, 1863
 * Frullania gigantea Stephani, 1911
 * Frullania giraldiana Massalongo, 1897
 * Frullania giraldiana var. handelii (Verdoorn) Hattori, 1972
 * Frullania globosa Hattori & Streimann, 1985
 * Frullania glomerata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg Ex Montagne, 1838
 * Frullania gracilicaulis Hattori, 1977
 * Frullania gracilis Subsp. (Gracilis 1978) (Reinwardt, Blume & Nees) Gottsche In G., L. & N., 1845
 * Frullania gracilis Subsp. (Gracilis 1978) var. brevior Nees In G., L. & N., 1845
 * Frullania gracilis Subsp. (Gracilis 1978) var. vittata Hattori, 1986
 * Frullania gracilis Subsp. Zennoskei Hattori & Thaithong, 1978
 * Frullania gradsteinii Yuzawa, Mues & Hattori, 1987
 * Frullania granatensis Gottsche, 1864
 * Frullania grandifolia Stephani, 1911
 * Frullania grandilobula Hattori & Piippo, 1986
 * Frullania grandistipula Lindenberg In G., L. & N., 1845
 * Frullania griffithsiana Gottsche In G., L. & N., 1846
 * Frullania grolleana Hattori, 1972
 * Frullania grossiclava Stephani, 1910
 * Frullania grossifolia Stephani, 1911
 * Frullania guadalupensis Gottsche Ex Stephani, 1911
 * Frullania gualaquizana Stephani, 1911
 * Frullania guatemalensis Stephani, 1911

H
 * Frullania haeckeriana Lindenberg In G., L. & N., 1846
 * Frullania haematocysta Spruce, 1884
 * Frullania hainanensis Hattori & Lin, 1986
 * Frullania hamata Stephani, 1911
 * Frullania hamaticoma Stephani, 1889
 * Frullania hamatiloba Stephani, 1910
 * Frullania hamatiloba F. Grosse-Appendiculata Hattori, 1944
 * Frullania hamatiloba var. latistipula Hattori, 1975
 * Frullania hamatosetacea Grolle In Grolle & Meister, 2004
 * Frullania hamiflora Herzog & Clark In Clark & Schultz, 1953
 * Frullania handelii Verdoorn In Handel-Mazzetti, 1930
 * Frullania handel-Mazzettii Hattori, 1981
 * Frullania hariotana Stephani, 1911
 * Frullania harpantha Herzog, 1942
 * Frullania hasskarliana Lindenberg In G., L. & N., 1845
 * Frullania hasskarliana var. gracilis Hattori, 1986
 * Frullania hasskarliana var. integribractea (Verdoorn) Hattori, 1975
 * Frullania hasskarliana var. parvidentata Hattori, 1986
 * Frullania hattoriana Godfrey & Godfrey, 1980
 * Frullania hattoriantha Udar & Nath, 1981
 * Frullania hattorii Konrat & Braggins, 2003
 * Frullania hawaiiensis Miller, 1953
 * Frullania hebridensis Stephani, 1911
 * Frullania hedrantha Hattori & Kamimura, 1973
 * Frullania helleri Stephani, 1910
 * Frullania herzogiana Stephani, 1911
 * Frullania herzogii Hattori, 1955
 * Frullania heteromorpha Schiffner, 1889
 * Frullania hicksiae Hattori, 1984
 * Frullania hicksiae F. Litoralis Hattori, 1988
 * Frullania higuchii Yuzawa & Hattori In Yuzawa & Koike, 1994
 * Frullania himalayensis Stephani, 1910
 * Frullania hinoi Kamimura, 1982
 * Frullania hiroshii Hattori, 1980
 * Frullania hirtiflora Spruce, 1884
 * Frullania holostipula Hattori & Griffin Iii, 1978
 * Frullania hottana Hattori, 1976
 * Frullania howeana Stephani In Stephani & Watts, 1914
 * Frullania huerlimannii Hattori, 1976
 * Frullania huerlimannii var. dioica Hattori, 1984
 * Frullania humbertii Vanden Berghen, 1976
 * Frullania humilis Spruce, 1890
 * Frullania hypoleuca Nees In G., L. & N., 1843
 * Frullania hypoleucula Hattori, 1984

I
 * Frullania imerinensis Stephani, 1911
 * Frullania immersa Stephani, 1896
 * Frullania incisoduthiana Hattori In Mizutani, 1979
 * Frullania incisoduthiana var. parva Hattori In Mizutani, 1979
 * Frullania incisostipula Stephani, 1924
 * Frullania inconstans Verdoorn, 1930
 * Frullania inconstans F. Integrior Verdoorn, 1930
 * Frullania inconstans var. grossedentata Kamimura & Hattori, 1973
 * Frullania incumbens Mitten In Hooker, 1855
 * Frullania incurva Hattori, 1988
 * Frullania inflata Gottsche In G., L. & N., 1845
 * Frullania inflata var. communis Schuster, 1985
 * Frullania inflata var. dioica Hattori & Thaithong, 1978
 * Frullania inflexa Mitten, 1861
 * Frullania inflexiloba Hattori, 1984
 * Frullania inouei Hattori, 1980
 * Frullania integristipula (Nees) Nees In G., L. & N., 1845
 * Frullania integristipula var. emarginata Verdoorn, 1929
 * Frullania intermedia Subsp. (Intermedia 1980) F. Billardieriana (Nees & Montagne In Montagne) Verdoorn, 1930
 * Frullania intermedia Subsp. (Intermedia 1980) var. non-Apiculata Hattori, 1975
 * Frullania intermedia Subsp. (Intermedia) (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
 * Frullania intermedia Subsp. Morokensis (Stephani) Hattori, 1980
 * Frullania intermixta Colenso, 1889
 * Frullania intumescens (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania involuta Hampe Ex Stephani, 1911
 * Frullania involvens Hattori & Kamimura, 1973
 * Frullania iriomotensis Hattori, 1980
 * Frullania irregularis Hattori & Piippo, 1986
 * Frullania iwatsukii Hattori, 1972

J
 * Frullania jackii Subsp. (Jackii 1959) Gottsche In Gottsche & Rabenhorst, 1863
 * Frullania jackii Subsp. (Jackii 1959) F. Depauperata Grolle, 1970
 * Frullania jacobsii Hattori, 1986
 * Frullania jacquinotii Gottsche Ex Stephani, 1910
 * Frullania jelskii Loitlesberger, 1894
 * Frullania johnsonii Stephani, 1894
 * Frullania junghuhniana Gottsche In G., L. & N., 1845
 * Frullania junghuhniana var. bisexualis Hattori, 1976
 * Frullania junghuhniana var. tenella (Sande Lacoste) Grolle & Hattori In Hattori, 1982
 * Frullania junghuhniana var. tenella F. Monoica (Hattori) Hattori, 1986

K
 * Frullania kagoshimensis Subsp. (Kagoshimensis 1985) Stephani, 1910
 * Frullania kagoshimensis Subsp. (Kagoshimensis 1985) var. minor Kamimura, 1961
 * Frullania kagoshimensis Subsp. Hunanensis (Hattori) Hattori, 1985
 * Frullania kalimantanensis Hattori, 1986
 * Frullania kalimantanensis Piippo & Hattori In Piippo & Tan, 1992
 * Frullania kashyapii Verdoorn, 1932
 * Frullania kitagawana Hattori, 1984
 * Frullania klotzschii Nees Ex Stephani, 1911
 * Frullania koponenii Hattori, 1978
 * Frullania kunzei (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania kunzei var. maritima Schuster, 1991

L
 * Frullania laetevirens Hampe In G., L. & N., 1845
 * Frullania laevi-Periantha Bai & Gao, 2000
 * Frullania lancistyla Stephani, 1910
 * Frullania larjiana Singh & Singh, 2005
 * Frullania laticaulis Spruce, 1890
 * Frullania latiflora Spruce, 1884
 * Frullania latogaleata Herzog, 1948
 * Frullania laxiflora Spruce, 1884
 * Frullania laxiflora var. crossii Spruce, 1884
 * Frullania leana Austin, 1869
 * Frullania leeuwenii Verdoorn, 1930
 * Frullania lepida Hattori & Piippo, 1986
 * Frullania letestui Vanden Berghen, 1976
 * Frullania levieri Stephani, 1910
 * Frullania libera Nees In G., L. & N., 1845
 * Frullania lindbergiana Gottsche, 1863
 * Frullania lindenbergii Gottsche Ex Lehmann, 1844
 * Frullania lindenbergii var. (Beta ?) Fusca Gottsche In G., L. & N., 1847
 * Frullania lindeniana Stephani, 1911
 * Frullania lindmanii Stephani, 1897
 * Frullania linii Hattori, 1981
 * Frullania lobato-Hastata Stephani, 1911
 * Frullania lobulata (Hooker) Hooker & Nees In G., L. & N., 1845
 * Frullania longipinna Stephani, 1910
 * Frullania longistipula Stephani In Renauld & Cardot, 1891
 * Frullania longistipula var. apiculata Demaret & Vanden Berghen, 1950
 * Frullania longistyla Yuzawa & Hattori, 1988
 * Frullania loricata Pearson, 1891
 * Frullania loricata var. laxa Pearson, 1891
 * Frullania ludoviciae Stephani, 1908
 * Frullania lushanensis Hattori & Lin, 1985

M
 * Frullania macgregorii Stephani, 1894
 * Frullania macgregorii var. rostellula (Hattori) Hattori, 1982
 * Frullania macrocephala (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania macrophylla Hattori, 1980
 * Frullania macularis Taylor, 1846
 * Frullania madagascariensis Gottsche, 1882
 * Frullania madens Stephani, 1924
 * Frullania madothecoides Spruce, 1884
 * Frullania magellanica (Sprengel) Weber & Nees In G., L. & N., 1845
 * Frullania magellanica var. diminutiva Herzog, 1938
 * Frullania mammilligera Grolle, 2003
 * Frullania mammillosa Hattori, 1977
 * Frullania matafaoica Miller, 1981
 * Frullania mathanii Stephani, 1911
 * Frullania mauritiana Austin, 1869
 * Frullania maymyoensis Svihla, 1958
 * Frullania mcveanii Hattori, 1973
 * Frullania media (Hodgson) Hattori, 1983
 * Frullania megalostipa Spruce, 1884
 * Frullania meijeri Hattori, 1974
 * Frullania meridana Stephani, 1911
 * Frullania meyeniana Lindenberg In G., L. & N., 1845
 * Frullania meyeniana var. dioica Hattori, 1977
 * Frullania microauriculata Verdoorn, 1929
 * Frullania microauriculata var. rotundior Verdoorn, 1929
 * Frullania microcaulis Gola, 1923
 * Frullania microcephala Gottsche, 1863
 * Frullania microphylla (Gottsche In Gottsche & Rabenhorst) Pearson, 1894
 * Frullania microphylla F. Corticicola Schiffner, 1936
 * Frullania microphylla var. deciduifolia Grolle, 1970
 * Frullania microrhyncha Clark & Svihla, 1950
 * Frullania microscopica Pearson, 1922
 * Frullania minor Subsp. (Minor 1975) var. integribracteola Hattori, 1975
 * Frullania minor Subsp. Recurviloba Hattori, 1975
 * Frullania mirabilis Jack & Stephani In Stephani, 1892
 * Frullania miradorensis Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania mizutanii Kamimura & Hattori, 1973
 * Frullania moniliata Subsp. Breviramea (Stephani) Verdoorn, 1930
 * Frullania moniliata Subsp. Moniliata Crandall-Stotler, Geissler & Stotler, 1987
 * Frullania moniliata Subsp. Moniliata F. Minshanensis (Hattori) Crandall-Stotler Et Alii, 1987
 * Frullania moniliata Subsp. Moniliata var. elongatistipula (Verdoorn) Crandall-Stotler Et Alii, 1987
 * Frullania moniliata Subsp. Moniliata var. vietnamica (Hattori) Crandall-Stotler Et Alii, 1987
 * Frullania monocera (Hooker & Taylor) Taylor In G., L. & N., 1845
 * Frullania monocera var. depauperata Hattori, 1984
 * Frullania monocera var. schiffneri (Verdoorn) Hattori, 1979
 * Frullania monoica Stephani, 1900
 * Frullania montagnei Gottsche In G., L. & N., 1845
 * Frullania montana Stephani, 1910
 * Frullania moritziana Lindenberg & Gottsche In G., L. & N, 1847
 * Frullania moritziana var. (Beta) Mexicana Gottsche, 1863
 * Frullania morobensis Hattori & Streimann, 1985
 * Frullania motoyana Stephani, 1911
 * Frullania mucronata (Lehmann & Lindenberg In Lehmann) Lehmann & Linde,nberg
 * Frullania mucronata var. submutica Lehmann & Lindenberg In G., L. & N. 1845 In G., L. & N., 1845
 * Frullania multilacera Subsp. (Multilacera 1987) Stephani, 1911
 * Frullania multilacera Subsp. (Multilacera 1987) var. lacerissima Hattori, 1975
 * Frullania multilacera Subsp. Gracilior Hattori, 1987
 * Frullania multilaceroides Hattori, 1987
 * Frullania muscicola Stephani, 1894
 * Frullania mutilata Stephani, 1911

N
 * Frullania nadeaudii Stephani, 1911
 * Frullania neocaledonica Engel In Engel & Merrill, 1999
 * Frullania neosheana Hattori, 1979
 * Frullania nepalensis (Sprengel) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania neurota Taylor, 1846
 * Frullania nicholsonii Hodgson, 1949
 * Frullania nigricaulis (R., B. & N.) Nees In G., L. & N., 1845
 * Frullania nigricaulis var. elongata Verdoorn, 1930
 * Frullania nisquallensis Sullivant, 1849
 * Frullania nivimontana Hattori, 1982
 * Frullania nobilis Stephani, 1894
 * Frullania nobilis var. cochleata (Stephani) Hattori In Hattori & Streimann, 1985
 * Frullania nodulosa (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
 * Frullania notarsii Stephani, 1911
 * Frullania novocurvirostris Hattori, 1981
 * Frullania novoguineensis Schiffner, 1890

O
 * Frullania oahuensis Hampe In G., L. & N., 1843
 * Frullania oakesiana Subsp. (Oakesiana 1992) Austin, 1870
 * Frullania oakesiana Subsp. Takayuensis (Stephani) Schuster, 1992
 * Frullania obovata Hattori, 1982
 * Frullania obscurifolia Mitten, 1879
 * Frullania ocanniensis Stephani, 1924
 * Frullania ocellata Kamimura & Hattori, 1973
 * Frullania odontostipa Spruce, 1890
 * Frullania okinawensis Kamimura, 1982
 * Frullania onraedtii Vanden Berghen, 1976
 * Frullania orbicularis Austin, 1869
 * Frullania orientalis Sande Lacoste, 1855
 * Frullania orinocensis Spruce, 1883
 * Frullania ornithocephala (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
 * Frullania ornithocephala F. Magnilobula Hattori, 1976
 * Frullania ornithocephala F. Retusa Hattori, 1976
 * Frullania ornithocephala F. Teres (Sande Lacoste) Verdoorn, 1929
 * Frullania ornithocephala var. maior Nees, 1845
 * Frullania ornithocephala var. minor Nees, 1845
 * Frullania ornithocephala var. pilosa Verdoorn, 1929
 * Frullania ornithocephala var. tuberculosa Hattori, 1974
 * Frullania osculatiana Notaris, 1855
 * Frullania osumiensis (Hattori) Hattori In Iwatsuki & Hattori, 1956
 * Frullania osumiensis var. dentata Kamimura, 1970

P
 * Frullania pachyderma Hattori, 1978
 * Frullania pallide-Virens Stephani, 1911
 * Frullania pallidula Hattori, 1988
 * Frullania pancheri Gottsche Ex Stephani, 1894
 * Frullania papillata Stephani, 1911
 * Frullania papillilobula Hattori, 1975
 * Frullania papuana Verdoorn, 1930
 * Frullania papulosa Stephani, 1911
 * Frullania paradoxa Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania paradoxa F. Cornistipula Herzog, 1952
 * Frullania parhami (Schuster) Schuster, 1992
 * Frullania pariharii Hattori & Thaithong, 1978
 * Frullania pariharii F. Intermedia Hattori & Thaithong, 1978
 * Frullania parvifolia Stephani, 1910
 * Frullania parvistipula Stephani, 1910
 * Frullania patagonica Stephani, 1910
 * Frullania patula Mitten In Hooker, 1855
 * Frullania pauciramea Stephani, 1911
 * Frullania pauciramea var. pauciramella Hattori & Piippo, 1986
 * Frullania paucirameoides Hattori & Piippo, 1986
 * Frullania pearceana Stephani, 1911
 * Frullania pedicellata Stephani, 1897
 * Frullania pentapleura Subsp. (Pentapleura 1988) Taylor, 1846
 * Frullania peruviana Gottsche In G., L. & N., 1846
 * Frullania phalangiflora Stephani, 1916
 * Frullania philippinensis Stephani, 1911
 * Frullania physantha Mitten Mitten, 1861
 * Frullania pilibracteola Hattori, 1977
 * Frullania piliflora Stephani, 1911
 * Frullania piliflora var. appendiculata Herzog, 1951
 * Frullania pilistipula Stephani, 1911
 * Frullania piptophylla Hattori, 1980
 * Frullania piptophylla var. minor Hattori, 1986
 * Frullania piptophylloides Hattori, 1980
 * Frullania pittier I Stephani, 1892
 * Frullania plana Sullivant, 1849
 * Frullania planifolia Stephani, 1910
 * Frullania platycalyx Herzog, 1952
 * Frullania platyphylla Colenso, 1889
 * Frullania pluricarinata Gottsche, 1864
 * Frullania pocsantha Thaithong & Hattori, 1977
 * Frullania polyclada Colenso, 1889
 * Frullania polyptera Taylor, 1846
 * Frullania polyptera var. angustata (Mitten) Hattori, 1975
 * Frullania polysticta Lindenberg In G., L. & N., 1845
 * Frullania ponapensis Hattori & Koike In Koike, 1994
 * Frullania pringlei Fulford & Sharp, 1990
 * Frullania probosciphora Taylor, 1846
 * Frullania prominula Hattori & Streimann, 1985
 * Frullania propaginea Hattori & Streimann, 1985
 * Frullania pseudericoides Hattori, 1982
 * Frullania pseudericoides Hattori, 1986
 * Frullania pseudomeyeniana Hattori, 1986
 * Frullania pseudomonocera Hattori, 1986
 * Frullania pseudoschensiana Hattori, 1980
 * Frullania pseudoschensiana var. darjeelingensis Hattori, 1981
 * Frullania ptychantha Montagne, 1843
 * Frullania pulchella Herzog, 1954
 * Frullania pullei Verdoorn, 1930
 * Frullania pulogensis Stephani, 1911
 * Frullania punctata Reimers, 1931
 * Frullania purpurea Stephani, 1911
 * Frullania pusilla Mitten, 1871
 * Frullania pycnantha (Hooker & Taylor) Taylor In G., L. & N., 1845
 * Frullania pycnoclada Grolle In Grolle & Meister, 2004
 * Frullania pyricalycina Stephani, 1910

Q
 * Frullania queenslandica Stephani, 1910
 * Frullania quillotensis (Nees & Montagne) Nees & Montagne In Montagne In Alcide D'orbigny, 1839
 * Frullania quillotensis F. Flaccida Herzog, 1954
 * Frullania quillotensis F. Leiantha Herzog, 1954

R
 * Frullania rabenhorstii Stephani, 1911
 * Frullania ramuligera (Nees) Montagne, 1842
 * Frullania recurvistipula Hattori, 1975
 * Frullania reflexistipula Sande Lacoste, 1853
 * Frullania reflexistipula var. novocaledonica Hattori, 1977
 * Frullania reflexistipula var. squarrosa Hattori & Piippo, 1986
 * Frullania regularis Schiffner, 1890
 * Frullania reicheana Stephani, 1910
 * Frullania reimersii Verdoorn, 1930
 * Frullania remotidens Hattori, 1872
 * Frullania remotifolia Stephani, 1911
 * Frullania remotiloba Stephani, 1894
 * Frullania repanda Gottsche, 1864
 * Frullania repandistipula Subsp. (Repandistipula 1975) Sande Lacoste, 1853
 * Frullania repandistipula Subsp. Queenslandica Hattori, 1987
 * Frullania repandistipula Subsp. Spinibractea Hattori, 1975
 * Frullania reptans var. integristipula Nicholson, 1925
 * Frullania retusa Mitten, 1861
 * Frullania retusa var. gymnantha Hattori & Thaithong, 1978
 * Frullania retusa var. hirsuta Hattori & Thaithong, 1978
 * Frullania rhystocolea Herzog Ex Verdoorn In Handel-Mazzetti, 1930
 * Frullania rhytidantha Hattori, 1980
 * Frullania rigescens Spruce, 1884
 * Frullania rigida Stephani, 1910
 * Frullania ringens Spruce, 1884
 * Frullania riojaneirensis (Raddi) Ångström, 1876
 * Frullania riparia Hampe Ex Lehmann, 1838
 * Frullania rizalli Piippo & Hattori, 1992
 * Frullania rostellata Mitten In Hooker, 1867
 * Frullania rostrata (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
 * Frullania rubella Gottsche Ex Stephani, 1889
 * Frullania rubella var. elongata (Stephani) Hattori, 1983
 * Frullania rudolfiana Hattori, 1972
 * Frullania rupicola Stephani, 1924

S
 * Frullania sabahana Hattori, 1976
 * Frullania sabaliana Schuster, 1983
 * Frullania sabanetica Gottsche, 1864
 * Frullania sachapatensis Stephani, 1911
 * Frullania sackawana Stephani, 1897
 * Frullania sackawana var. minor Kamimura, 1982
 * Frullania saipanensis Hattori & Koike In Koike, 1994
 * Frullania sarawakensis Hattori, 1976
 * Frullania scalaris Hattori, 1977
 * Frullania scandens Montagne, 1843
 * Frullania schaefer-Verwimpii Yuzawa & Hattori, 1989
 * Frullania schensiana Massalongo, 1897
 * Frullania schiffneri Verdoorn, 1929
 * Frullania schimperi Nees In G., L. & N., 1845
 * Frullania schimperi var. laciniata Vanden Berghen, 1976
 * Frullania schumannii (Caspary) Grolle, 1981
 * Frullania schusteri Hattori, 1988
 * Frullania schusteriana Hattori, 1972
 * Frullania scottiana Hattori, 1987
 * Frullania sebastianopolitana var. (Delta) Gottsche, 1864
 * Frullania selwyniana Pearson, 1890
 * Frullania semienana Gola, 1914
 * Frullania semivillosa Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania semivillosa var. (Beta) Glabra Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania sergiae Sim-Sim, Fontihna, Mues & Lion, 2000
 * Frullania seriata Gottsche Ex Stephani, 1889
 * Frullania seriatifolia Stephani, 1894
 * Frullania serrata Subsp. (Serrata 1982) Gottsche In G., L. & N., 1845
 * Frullania serrata Subsp. (Serrata 1982) F. Crispulodentata Verdoorn, 1929
 * Frullania serrata Subsp. (Serrata 1982) F. Vogelkopensis Hattori, 1982
 * Frullania serrata Subsp. (Serrata 1982) var. ceramensis Hattori, 1986
 * Frullania serrata Subsp. (Serrata 1982) var. hamatispina (Hattori) Hattori, 1982
 * Frullania serrata Subsp. (Serrata 1982) var. oceanica Verdoorn, 1937
 * Frullania serrata Subsp. (Serrata 1982) var. pertenuis (Nees) Gottsche In G., L. & N., 1845
 * Frullania serrata Subsp. Grolleana Grolle & Piippo, 1984
 * Frullania serrata Subsp. Spinistipula (Hattori) Hattori, 1982
 * Frullania setacea Hattori, 1988
 * Frullania setigera Stephani, 1894
 * Frullania shanensis Svihla, 1957
 * Frullania sharpantha Udar & Kumar, 1983
 * Frullania sharpii Hattori, 1974
 * Frullania sheana Hattori, 1979
 * Frullania simmondsii Stephani In Stephani & Watts, 1914
 * Frullania sinensis Stephani In Levier, 1906
 * Frullania sinosphaerantha Hattori & Lin, 1985
 * Frullania sinuata Subsp. Sinuata 1975 Sande Lacoste, 1853
 * Frullania sinuata Subsp. Sinuata F. Parvistylata Hattori In Grolle & Piippo, 1984
 * Frullania socotrana Mitten In Balfour, 1888
 * Frullania solanderiana Colenso, 1889
 * Frullania spathulistipa Stephani, 1910
 * Frullania speciosa Herzog, 1949
 * Frullania spegazzinii Reiner, 1988
 * Frullania sphaerantha Hattori, 1980
 * Frullania sphaerocephala Spruce, 1884
 * Frullania sphaerolobulata Lin In Lin & Chen, 1997
 * Frullania spicata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania spinifera Taylor, 1846
 * Frullania spinigastria Hattori, 1979
 * Frullania spiniplica Hattori, 1972
 * Frullania spinistipula Stephani, 1911
 * Frullania spongiosa Stephani, 1894
 * Frullania squamuligera Spruce, 1884
 * Frullania squarrosula (Hooker & Taylor) Taylor In G., L. & N., 1845
 * Frullania standaerti Stephani, 1910
 * Frullania steereana Hattori, 1987
 * Frullania stenostipa Spruce, 1884
 * Frullania streimannii Hattori, 1983
 * Frullania stylifera (Schuster) Schuster, 1992
 * Frullania subcaduca Hattori, 1974
 * Frullania subclavata Stephani, 1910
 * Frullania subdentata Stephani, 1911
 * Frullania subdentata F. Longistipula Hattori, 1978
 * Frullania subdilatata Massalongo In Levier, 1906
 * Frullania subhampeana Hodgson, 1949
 * Frullania subincumbens Hattori, 1987
 * Frullania sublignosa Stephani, 1894
 * Frullania submultilacera Hattori & Koike In Koike, 1994
 * Frullania subnigricaulis Hattori, 1973
 * Frullania subnigricaulis var. subtruncata Hattori, 1975
 * Frullania subocellata Hattori, 1986
 * Frullania subpedicellata Hattori, 1980
 * Frullania subpilibracteola Hattori, 1977
 * Frullania subpyricalycina Herzog, 1954
 * Frullania subrostrata Hattori, 1972
 * Frullania subsquarrosa Hattori, 1972
 * Frullania subtilissima Lindenberg In G., L. & N., 1845
 * Frullania subtropica Stephani, 1910
 * Frullania subtruncata Stephani, 1910
 * Frullania subvalida Hattori & Thaithong, 1978
 * Frullania supradecomposita (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
 * Frullania svihlana Hattori, 1983

T
 * Frullania tagawana (Hattori, Thaithong & Kitawana) Hattori In Hattori & Lin, 1985
 * Frullania taiheizana Horikawa, 1934
 * Frullania tamarisci (L.) Dumortier, 1835
 * Frullania tamarisci var. atlantica Schiffner, 1936
 * Frullania tamarisci var. atrovirens Carrington, 1863
 * Frullania tamarisci var. commutata Nees In G., L. & N., 1845
 * Frullania tamarisci var. cornubica Carrington In Carrington & Pearson, 1878
 * Frullania tamarisci var. laxa Nees In G., L. & N., 1845
 * Frullania tamarisci var. mediterranea (De Notaris) Nees In G., L. & N., 1845
 * Frullania tamarisci var. robusta Lindberg, 1875
 * Frullania tamarisci var. sardoa (De Notaris) De Notaris, 1865
 * Frullania tamarisci var. schiffneri Nicholson, 1933
 * Frullania tamsuina Stephani, 1910
 * Frullania taradakenis Stephani, 1910
 * Frullania taxodiocola Schuster, 1983
 * Frullania teneriffae (Weber) Nees, 1838
 * Frullania teneriffae F. Explanata (Macvicar) Bisang, Schumacker, Sérgio & Grolle, 1988
 * Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle, 1988
 * Frullania tenuirostris Stephani, 1911
 * Frullania ternatensis Gottsche In G., L. & N., 1846
 * Frullania ternatensis var. non-Appendiculata Hattori, 1974
 * Frullania tetraptera Nees & Montagne In Montagne, 1838
 * Frullania thiersiae Hattori, 1988
 * Frullania tixieri Hattori, 1976
 * Frullania tjibodensis Hattori & Thaithong, 1977
 * Frullania togashiana Hattori, 1975
 * Frullania tongariroense Colenso, 1889
 * Frullania trianae Gottsche, 1864
 * Frullania tricarinata Sande Lacoste, 1856
 * Frullania trichodes Mitten, 1862
 * Frullania trigona Clark, Jovet-Ast & Frye, 1947
 * Frullania trinervis (Lehmann & Lindenberg) Drège, 1843
 * Frullania triquetra Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania trollii Herzog, 1942
 * Frullania truncata Caspary, 1887
 * Frullania truncatistyla von Konrat et al., 2011
 * Frullania tubercularis Hattori & Lin, 1985
 * Frullania tunguraguana Clark & Frye, 1952
 * Frullania turfosa Lindenberg & Gottsche In G., L. & N., 1847
 * Frullania tuyamae Hattori & Thaithong, 1978

U
 * Frullania udarii Nath & Singh, 2006
 * Frullania umbonata Mitten Ex Stephani, 1911
 * Frullania undulata Kamimura, 1961
 * Frullania usambarana Schiffner Ex Stephani, 1894
 * Frullania usambarana var. reducta Vanden Berghen, 1976
 * Frullania usamiensis Stephani, 1897
 * Frullania utriculata Stephani, 1894

V
 * Frullania vaga Mitten In Seemann, 1871
 * Frullania vaginata (Swartz) Nees In G., L. & N., 1846
 * Frullania valdiviensis Jack & Stephani In Stephani, 1894
 * Frullania valida Stephani, 1910
 * Frullania valparaisiana Lehmann, 1857
 * Frullania vandenberghenii Pocs In Bizot & Pocs, 1979
 * Frullania van-Zantenii Kamimura & Hattori, 1973
 * Frullania variabilis Stephani, 1910
 * Frullania varians Caspary, 1887
 * Frullania variegata Stephani, 1894
 * Frullania venusta Hattori, 1974
 * Frullania verdoorniana Hattori, 1973
 * Frullania victoriensis Stephani, 1910
 * Frullania vitalii Yuzawa & Hattori, 1988
 * Frullania vittata Hattori, 1974
 * Frullania vittata F. Denticulata Hattori, 1980
 * Frullania vittiana Hattori, 1987

W
 * Frullania wagneri Stephani, 1910
 * Frullania wairua Konrat & Braggins, 2005
 * Frullania wangii Hattori & Lin, 1985
 * Frullania warnckeana Hattori, 1974
 * Frullania warnckeana var. dentosa Hattori, 1977
 * Frullania weberbaueri Stephani, 1911
 * Frullania winteri Stephani, 1910
 * Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, 1991

Y
 * Frullania yorkiana Stephani, 1911
 * Frullania yuennanensis Stephani, 1894
 * Frullania yuennanensis var. siamensis (Hatt. et al.) Hattori & Lin, 1985
 * Frullania yusawae Kamimura, 1970
 * Frullania yuzawana Hattori, 1981

Z
 * Frullania zangii Hattori & Lin, 1985
 * Frullania zennoskeana Hattori, 1984

Extinct species

 * †Frullania baerlocheri Heinrichs et al., 2012 Burmese amber, Myanmar, Cenomanian
 * †Frullania cretacea Hentschel et al., 2009 Burmese amber, Myanmar, Cenomanian
 * †Frullania partita Li et al., 2020 Burmese amber, Myanmar, Cenomanian
 * †Frullania pinnata Heinrichs et al., 2017 Burmese amber, Myanmar, Cenomanian