Gomphillaceae

The Gomphillaceae are a family of lichen-forming fungi in the order Graphidales. Species in this family are found mostly in tropical regions.

Description
The family Gomphillaceae is characterized by its crust-like thallus, which is the outer layer of the lichen body. These lichens contain photobionts, which are symbiotic algae that help the lichen to produce food through photosynthesis. The reproductive structures of Gomphillaceae lichens, known as ascomata, can be apothecioid or lirellate in form. Apothecia occur in a few forms, including or  to sometimes.

The internal structure of hymenium (the fertile, spore-producing part of the lichen), called the, consists primarily of branched and interwoven. The asci, or spore-producing sacs, are and feature a distinctive apical  and ring structure. They can be, oblong, or in shape and do not exhibit any amyloid properties. These asci typically produce eight, although some may produce fewer, ranging from one to four spores. The ascospores are hyaline (transparent), and have thin walls and distinct. They can be ellipsoid or oblong in shape, and their internal divisions (septa) can be transverse or.

Gomphillaceae lichens also produce, which are asexual reproductive structures. These are mostly, and the conidia are formed as branched hyphae, called diahyphae, within gelatinous masses. The conidia are typically septate, often taking on a or bead-like appearance, and are also hyaline. In terms of secondary chemistry, Gomphillaceae lichens generally lack any notable substances.

Genera
According to a recent (2022) estimate, the Gomphillaceae comprise 26 genera and about 425 species. The following list indicates the genus name, the taxonomic authority, year of publication, and the number of species:
 * Actinoplaca Müll.Arg. (1891) – 2 spp.
 * Aderkomyces Bat. (1961) – 30 spp.
 * Aplanocalenia Lücking, Sérus. & Vězda (2005) – 1 sp.
 * Arthotheliopsis Vain. (1896) – 5 spp.
 * Asterothyrium Müll.Arg. (1890) – 32 spp.
 * Aulaxina Fée (1825) – 14 spp.
 * Calenia Müll.Arg. (1890) – 30 spp.
 * Caleniopsis Vězda & Poelt (1987) – 2 spp.
 * Corticifraga D.Hawksw. & R.Sant. (1990) – 9 spp.
 * Diploschistella Vain. (1926) – 4 spp.
 * Echinoplaca Fée (1825) – 40 spp.
 * Ferraroa Lücking, Sérus. & Vězda (2005) – 1 sp.
 * Gomphillus Nyl. (1855) – 6 spp.
 * Gyalectidium Müll.Arg. (1881) – 52 spp.
 * Gyalidea Lettau ex Vězda (1996) – 50 spp.
 * Gyalideopsis Vězda (1972) – 91 spp.
 * Hippocrepidea Sérus. (1997) – 1 sp.
 * Jamesiella Lücking, Sérus. & Vězda (2005) – 4 spp.
 * Lithogyalideopsis Lücking, Sérus. & Vězda (2005) – 4 spp.
 * Paragyalideopsis Etayo (2017) – 4 spp.
 * Paratricharia Lücking (1997) – 1 spp.
 * Phyllogyalidea Lücking & Aptroot (2008) – 2 spp.
 * Psorotheciopsis Rehm (1900) – 7 spp.
 * Rolueckia Papong, Thammath. & Boonpr. (2008) – 3 spp.
 * Taitaia Suija, Kaasalainen, Kirika & Rikkinen (2018) – 1 spp.
 * Tricharia Fée (1825) – ca. 30 spp.