Harmonia (beetle)

Harmonia is a genus of lady beetles belonging to the subfamily Coccinellinae.

Description
The distinction of the individual species within the genus is often very difficult, and in some cases requires examination of the reproductive system. The coloring and drawing in the genus Harmonia are very variable, not only between species, but also within each species. The basic color of the elytra is usually red to orange with black spots. The patches can be linked together to form black bars. The species can reach a length of 3 to 7 mm. The rear edge of the pronotum is slightly curved. The antennae are a little shorter than the head and are composed of eleven segments. The three last segments are thickened at the tip and form a club.

List of species

 * Harmonia axyridis (Pallas, 1771) – Asian lady beetle, originally from Nepal, China, Taiwan, Japan, introduced to Western Europe and America
 * Harmonia antipodum (Mulsant, 1848) – antipodean ladybird, endemic to New Zealand
 * Harmonia basinotata Bielawski, 1964 – New Guinea
 * Harmonia bicolor (Blackburn, 1892) – Australia
 * Harmonia conformis (Boisduval, 1835) – common spotted lady beetle, Australia, introduced in New Zealand
 * Harmonia dimidiata (Fabricius, 1781) – India, Pakistan, Nepal, Bhutan, China, Taiwan, Japan; introduced in North America
 * Harmonia dunlopi (Crotch, 1874) – India, Malaysia, Philippines
 * Harmonia eucharis (Mulsant, 1850) – India, Pakistan, Myanmar, Nepal, Himalayan Region, South China
 * Harmonia expallida (Weise, 1907) – India
 * Harmonia octomaculata (Fabricius, 1781) – India, Pakistan, Nepal, Bangladesh, Sri Lanka, Micronesia, Australia
 * Harmonia quadripunctata (Pontoppidan, 1763) – cream-streaked lady beetle, Europe
 * Harmonia sedecimnotata (Fabricius, 1801) – India, Nepal, Southeast Asia
 * Harmonia testudinaria (Mulsant, 1850) – Indonesia, New Guinea, North Australia
 * Harmonia uninotata Bielawski, 1964 – New Guinea
 * Harmonia yedoensis (Takizawa, 1917) – Japan