Harpalus (beetle)

Harpalus is a genus of ground beetle first described by Pierre André Latreille in 1802.

Species
Harpalus contains the following 430 species:
 * Subgenus Cephalophonus Ganglbauer, 1891
 * Harpalus cephalotes Fairmaire & Laboulbène, 1854


 * Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982


 * Harpalus agnatus Reiche, 1850
 * Harpalus cyrenaicus Koch, 1939
 * Harpalus fulvus Dejean, 1829
 * Harpalus grilli Kataev, 2002
 * Harpalus idiotus Bates, 1889
 * Harpalus janinae Jeanne, 1984
 * Harpalus litigiosus Dejean, 1829
 * Harpalus melancholicus Dejean, 1829
 * Harpalus schaumii Wollaston, 1864
 * Harpalus tenebrosus Dejean, 1829


 * Subgenus Glanodes Casey, 1914
 * Harpalus cohni Ball, 1972
 * Harpalus corpulentus (Casey, 1914)
 * Harpalus huachuca Ball, 1972
 * Harpalus obliquus G.Horn, 1880
 * Harpalus puncticeps (Casey, 1914)
 * Harpalus stephani Ball, 1972


 * Subgenus Harpalus Latreille, 1802


 * Harpalus acupalpoides Reitter, 1900
 * Harpalus aeneipennis (Faldermann, 1836)
 * Harpalus aequicollis Motschulsky, 1844
 * Harpalus affinis (Schrank, 1781)
 * Harpalus agakhaniantzi (Mikhailov, 1972)
 * Harpalus akbensis Jedlicka, 1958
 * Harpalus akinini Tschitscherine, 1895
 * Harpalus alajensis Tschitscherine, 1898
 * Harpalus albanicus Reitter, 1900
 * Harpalus alexandrae Kataev, 1990
 * Harpalus alexeevi Kataev, 1990
 * Harpalus alpivagus Tschitscherine, 1899
 * Harpalus amarellus Bates, 1891
 * Harpalus amariformis Motschulsky, 1844
 * Harpalus amplicollis Ménétriés, 1848
 * Harpalus amputatus Say, 1830
 * Harpalus anatolicus Tschitscherine, 1898
 * Harpalus angulatus Putzeys, 1878
 * Harpalus angustipennis Boheman, 1848
 * Harpalus angustitarsis Reitter, 1887
 * Harpalus animosus Casey, 1924
 * Harpalus anisodactyliformis Solsky, 1874
 * Harpalus antonowi (Tschitscherine, 1898)
 * Harpalus anxioides Kataev, 1991
 * Harpalus anxius (Duftschmid, 1812)
 * Harpalus araraticus Mlynar, 1979
 * Harpalus arcuatus Tschitscherine, 1898
 * Harpalus arnoldii Kataev, 1988
 * Harpalus ascetes Kataev & Wrase, 1997
 * Harpalus asemus Basilewsky, 1946
 * Harpalus aterrimus Casey, 1914
 * Harpalus atratus Latreille, 1804
 * Harpalus atrichatus Hatch, 1949
 * Harpalus atripes Casey, 1914
 * Harpalus attenuatus Stephens, 1828
 * Harpalus australasiae Dejean, 1829
 * Harpalus autumnalis (Duftschmid, 1812)
 * Harpalus badakschanus Jedlicka, 1956
 * Harpalus baleensis Clarke, 1973
 * Harpalus balli Noonan, 1991
 * Harpalus basanicus J.Sahlberg, 1913
 * Harpalus basilewskyi Facchini, 2003
 * Harpalus basuto Basilewsky, 1958
 * Harpalus bellieri Reiche, 1861
 * Harpalus beneshi Kataev & Wrase, 1997
 * Harpalus brachypterus Tschitscherine, 1898
 * Harpalus brevicornis Germar, 1823
 * Harpalus brevis Motschulsky, 1844
 * Harpalus breviusculus Chaudoir, 1846
 * Harpalus bucharicus Tschitscherine, 1898
 * Harpalus bulirschi (Facchini, 2012)
 * Harpalus bungii Chaudoir, 1844
 * Harpalus caeruleatus Bates, 1878
 * Harpalus caiphus Reiche & Saulcy, 1855
 * Harpalus calathoides Motschulsky, 1844
 * Harpalus capicola Dejean, 1829
 * Harpalus cardoni Antoine, 1922
 * Harpalus caspius (Steven, 1806)
 * Harpalus cautus Dejean, 1829
 * Harpalus chalcentus Bates, 1873
 * Harpalus chasanensis Lafer, 1989
 * Harpalus chobautianus Lutshnik, 1922
 * Harpalus chrysopus Reitter, 1887
 * Harpalus circumpunctatus Chaudoir, 1846
 * Harpalus cisteloides Motschulsky, 1844
 * Harpalus clarkei Kataev & J.Schmidt, 2020
 * Harpalus compressus Motschulsky, 1844
 * Harpalus contemptus Dejean, 1829
 * Harpalus convexus Faldermann, 1836
 * Harpalus corporosus (Motschulsky, 1862)
 * Harpalus corrugatus Basilewsky, 1958
 * Harpalus crates Bates, 1883
 * Harpalus cupreus Dejean, 1829
 * Harpalus cursorius Péringuey, 1896
 * Harpalus cyanopterus (Tschitscherine, 1897)
 * Harpalus cyclogonus Chaudoir, 1844
 * Harpalus danieli Reitter, 1900
 * Harpalus davidianus Tschitscherine in Jakobson, 1903
 * Harpalus decipiens Dejean, 1829
 * Harpalus defector Péringuey, 1896
 * Harpalus desertus LeConte, 1859
 * Harpalus diligens Tschitscherine, 1898
 * Harpalus dimidiatus (P.Rossi, 1790)
 * Harpalus dispar Dejean, 1829
 * Harpalus dissitus Antoine, 1931
 * Harpalus distinguendus (Duftschmid, 1812)
 * Harpalus diversicollis Basilewsky, 1958
 * Harpalus dubius Boheman, 1848
 * Harpalus dudkoi Kataev, 2011
 * Harpalus durangoensis Bates, 1891
 * Harpalus ebeninus Heyden, 1870
 * Harpalus egenus Dejean, 1829
 * Harpalus egorovi Lafer, 1989
 * Harpalus ellipsis LeConte, 1847
 * Harpalus elliptipennis Facchini, 2003
 * Harpalus erosus Mannerheim, 1825
 * Harpalus exiguus Boheman, 1848
 * Harpalus famelicus Tschitscherine, 1898
 * Harpalus farkaci Kataev & Wrase, 1995
 * Harpalus fimetarius Dejean, 1829
 * Harpalus flavescens (Piller & Mitterpacher, 1783)
 * Harpalus flavicornis Dejean, 1829
 * Harpalus foveiger Tschitscherine, 1895
 * Harpalus franzi Mateu, 1954
 * Harpalus frater Chaudoir, 1876
 * Harpalus fraternus LeConte, 1852
 * Harpalus froelichii Sturm, 1818
 * Harpalus fulvicornis Thunberg, 1806
 * Harpalus fulvilabris Mannerheim, 1853
 * Harpalus fulvipennis Chaudoir, 1843
 * Harpalus fuscicornis Ménétriés, 1832
 * Harpalus fuscipalpis Sturm, 1818
 * Harpalus fuscipennis Wiedemann, 1825
 * Harpalus fuscoaeneus Dejean, 1829
 * Harpalus ganssuensis Semenov, 1889
 * Harpalus giacomazzoi Kataev & Wrase, 1996
 * Harpalus gilgil Basilewsky, 1946
 * Harpalus gisellae Csiki, 1932
 * Harpalus glasunovi Kataev, 1987
 * Harpalus gravis LeConte, 1858
 * Harpalus gregoryi Alluaud, 1917
 * Harpalus hartmanni Kataev, 2002
 * Harpalus herbivagus Say, 1823
 * Harpalus heyrovskyi Jedlicka, 1928
 * Harpalus hiekei Kataev & Wrase, 2010
 * Harpalus hirtipes (Panzer, 1796)
 * Harpalus honestus (Duftschmid, 1812)
 * Harpalus hospes Sturm, 1818
 * Harpalus hybridus Boheman, 1848
 * Harpalus impressus Roth, 1851
 * Harpalus inconcinnus Chaudoir, 1876
 * Harpalus indianus Csiki, 1932
 * Harpalus indicola Bates, 1878
 * Harpalus indigens Casey, 1924
 * Harpalus inexspectatus Kataev, 1989
 * Harpalus ingenuus Tschitscherine, 1898
 * Harpalus innocuus LeConte, 1863
 * Harpalus italus Schaum, 1860
 * Harpalus jeanneli Basilewsky, 1946
 * Harpalus jordanus Jedlicka, 1964
 * Harpalus kabakianus Kataev, 1988
 * Harpalus kabakovi Kataev, 1987
 * Harpalus kadleci Kataev & Wrase, 1995
 * Harpalus kadyrbekovi Kataev, 1988
 * Harpalus kagyzmanicus Kataev, 1984
 * Harpalus kandaharensis Jedlicka, 1956
 * Harpalus karakorum Jedlicka, 1958
 * Harpalus karamani Apfelbeck, 1902
 * Harpalus kazanensis Jedlicka, 1958
 * Harpalus kaznakovi Kataev & Wrase, 1997
 * Harpalus kibonoti Alluaud, 1927
 * Harpalus kirgisicus (Motschulsky, 1844)
 * Harpalus kiritshenkoi Kataev, 1990
 * Harpalus klapperichi Jedlicka, 1956
 * Harpalus kmecoi Facchini, 2003
 * Harpalus kozlovi Kataev, 1993
 * Harpalus kryzhanovskii Kataev, 1988
 * Harpalus kunarensis Kataev, 1993
 * Harpalus laetus Reiche, 1843
 * Harpalus laevipes Zetterstedt, 1828
 * Harpalus lama Kataev & Wrase, 1997
 * Harpalus lateralis Dejean, 1829
 * Harpalus laticeps LeConte, 1850
 * Harpalus latus (Linnaeus, 1758)
 * Harpalus lederi Tschitscherine, 1899
 * Harpalus lethierryi Reiche, 1860
 * Harpalus lewisii LeConte, 1865
 * Harpalus longipalmatus Mordkovitsh, 1969
 * Harpalus lopezi J.Serrano & Lencina, 2009
 * Harpalus lugubris Boheman, 1848
 * Harpalus lumbaris Mannerheim, 1825
 * Harpalus luteicornis (Duftschmid, 1812)
 * Harpalus lutshniki Schauberger, 1932
 * Harpalus macronotus Tschitscherine, 1893
 * Harpalus mairei Peyerimhoff, 1928
 * Harpalus major (Motschulsky, 1850)
 * Harpalus makhekensis Basilewsky, 1958
 * Harpalus manas Kataev, 1990
 * Harpalus marginellus Gyllenhal, 1827
 * Harpalus martini Van Dyke, 1926
 * Harpalus masoreoides Bates, 1878
 * Harpalus massarti Burgeon, 1936
 * Harpalus mauritanicus Gaubil, 1844
 * Harpalus medvedevi Kataev, 2006
 * Harpalus megacephalus LeConte, 1847
 * Harpalus melaneus Bates, 1878
 * Harpalus metallinus Ménétriés, 1836
 * Harpalus metarsius Andrewes, 1930
 * Harpalus meteorus Basilewsky, 1946
 * Harpalus michaili Kataev, 1990
 * Harpalus michailovi Kataev, 1987
 * Harpalus microthorax (Motschulsky, 1849)
 * Harpalus miles Péringuey, 1896
 * Harpalus minutissimus Facchini, 2015
 * Harpalus minutulus Kataev & Liang, 2004
 * Harpalus mitridati Pliginskiy, 1915
 * Harpalus mlynari Kataev, 1990
 * Harpalus modestus Dejean, 1829
 * Harpalus morvani Kataev, 2002
 * Harpalus murzini Kataev, 2008
 * Harpalus nanniscus Péringuey, 1896
 * Harpalus natalensis Boheman, 1848
 * Harpalus natalicus Péringuey, 1896
 * Harpalus neglectus Audinet-Serville, 1821
 * Harpalus nevadensis K. & J.Daniel, 1898
 * Harpalus nigrans A.Morawitz, 1862
 * Harpalus nigritarsis C.R.Sahlberg, 1827
 * Harpalus numidicus Bedel, 1893
 * Harpalus nyassicus Basilewsky, 1946
 * Harpalus oblitus Dejean, 1829
 * Harpalus obnixus Casey, 1924
 * Harpalus ochropus Kirby, 1837
 * Harpalus oodioides Dejean, 1829
 * Harpalus opacipennis (Haldeman, 1843)
 * Harpalus optabilis Dejean, 1829
 * Harpalus ovtshinnikovi Kataev, 1990
 * Harpalus pallidipennis A.Morawitz, 1862
 * Harpalus pallipes Chaudoir, 1837
 * Harpalus parallelocollis Facchini, 2003
 * Harpalus parasinuatus Kataev & Liang, 2007
 * Harpalus parvulus Dejean, 1829
 * Harpalus petri Tschitscherine, 1902
 * Harpalus picipennis (Duftschmid, 1812)
 * Harpalus plancyi Tschitscherine, 1897
 * Harpalus plenalis Casey, 1914
 * Harpalus politus Dejean, 1829
 * Harpalus polyglyptus Schaum, 1862
 * Harpalus potanini Tschitscherine, 1906
 * Harpalus praecurrens Schauberger, 1934
 * Harpalus procognatus Lorenz, 1998
 * Harpalus progrediens Schauberger, 1922
 * Harpalus prosperus Basilewsky, 1972
 * Harpalus providens Casey, 1914
 * Harpalus pseudoserripes Reitter, 1900
 * Harpalus pterostichus (Reitter, 1900)
 * Harpalus puetzi Kataev & Wrase, 1997
 * Harpalus pulchrinulus Reitter, 1900
 * Harpalus pulvinatus Ménétriés, 1848
 * Harpalus pumilus Sturm, 1818
 * Harpalus punctatostriatus Dejean, 1829
 * Harpalus punctipennis Mulsant, 1852
 * Harpalus pusillus (Motschulsky, 1850)
 * Harpalus pygmaeus Dejean, 1829
 * Harpalus quadratus Chaudoir, 1846
 * Harpalus raphaili Kataev, 1997
 * Harpalus reflexus Putzeys in Schneider & Leder, 1878
 * Harpalus reitteri Wrase & Kataev, 2011
 * Harpalus remboides Solsky, 1874
 * Harpalus reversus Casey, 1924
 * Harpalus rhemboides Solsky, 1874
 * Harpalus rivalsi Jeannel, 1948
 * Harpalus rotundus Facchini, 2003
 * Harpalus rougemonti Clarke, 1973
 * Harpalus rubripes (Duftschmid, 1812)
 * Harpalus rufipalpis Sturm, 1818
 * Harpalus rufiscapus Gebler, 1833
 * Harpalus rufitarsoides Schauberger, 1934
 * Harpalus rufomarginatus (Boheman, 1848)
 * Harpalus rumelicus Apfelbeck, 1904
 * Harpalus salinulus Reitter, 1900
 * Harpalus salinus Dejean, 1829
 * Harpalus sanctaehelenae Basilewsky, 1972
 * Harpalus saxicola Dejean, 1829
 * Harpalus semenowi Tschitscherine, 1901
 * Harpalus semipunctatus Dejean, 1829
 * Harpalus serripes (Quensel, 1806)
 * Harpalus servus (Duftschmid, 1812)
 * Harpalus setiporus (Reitter, 1894)
 * Harpalus siculus Dejean, 1829
 * Harpalus silipes Dejean, 1831
 * Harpalus sinuaticollis Facchini, 2015
 * Harpalus sinuatus Tschitscherine, 1893
 * Harpalus smaragdinus (Duftschmid, 1812)
 * Harpalus smyrnensis Heyden, 1888
 * Harpalus solitaris Dejean, 1829
 * Harpalus somereni Basilewsky, 1946
 * Harpalus somnulentus Dejean, 1829
 * Harpalus spadiceus Dejean, 1829
 * Harpalus spretus Péringuey, 1896
 * Harpalus spurius Péringuey, 1896
 * Harpalus stevenii Dejean, 1829
 * Harpalus stevensi Kataev, 2011
 * Harpalus stoetznerianus Schauberger, 1932
 * Harpalus strenuus Tschitscherine, 1898
 * Harpalus subaeneus Boheman, 1848
 * Harpalus subcylindricus Dejean, 1829
 * Harpalus subtruncatus Chaudoir, 1846
 * Harpalus sulphuripes Germar, 1823
 * Harpalus sundaicus Schauberger, 1933
 * Harpalus sushenicus Kataev, 1990
 * Harpalus suturangulus Reitter, 1887
 * Harpalus szalliesi Kataev & Wrase, 1995
 * Harpalus taciturnus Dejean, 1829
 * Harpalus tangutorum Kataev, 1993
 * Harpalus tardus (Panzer, 1796)
 * Harpalus tarsalis Mannerheim, 1825
 * Harpalus terrestris (Motschulsky, 1844)
 * Harpalus tibeticus Andrewes, 1930
 * Harpalus tichonis Jakobson, 1907
 * Harpalus tiridates Reitter, 1900
 * Harpalus tithonus Reitter, 1900
 * Harpalus tjanschanicus Semenov, 1889
 * Harpalus torosensis Jedlicka, 1961
 * Harpalus torridoides Reitter, 1900
 * Harpalus trichophorus Tschitscherine, 1897
 * Harpalus triseriatus A.Fleischer, 1897
 * Harpalus turcicus Jedlicka, 1958
 * Harpalus turmalinus Erichson, 1847
 * Harpalus udege Lafer, 1989
 * Harpalus uniformis (Motschulsky, 1844)
 * Harpalus ussuricus Mlynar, 1979
 * Harpalus vanemdeni Schauberger, 1932
 * Harpalus venator Boheman, 1848
 * Harpalus ventralis LeConte, 1847
 * Harpalus vereschaginae Kataev, 1988
 * Harpalus vernicosus Kataev & Liang, 2007
 * Harpalus viridanus (Motschulsky, 1844)
 * Harpalus viridicupreus Reiche, 1843
 * Harpalus vittatus Gebler, 1833
 * Harpalus wadiensis Jedlicka, 1964
 * Harpalus wohlberedti Emden, 1932
 * Harpalus xanthopus Gemminger & Harold, 1868
 * Harpalus zabroides Dejean, 1829
 * Harpalus zhdankoi Kataev, 1990


 * Subgenus Loboharpalus Schauberger, 1932
 * Harpalus platynotus Bates, 1873
 * Harpalus rubefactus Bates, 1873


 * Subgenus Opadius Casey, 1914
 * Harpalus apache Kataev, 2010
 * Harpalus cordatus (LeConte, 1853)
 * Harpalus cordifer Notman, 1919


 * Subgenus Plectralidus Casey, 1914
 * Harpalus erraticus Say, 1823
 * Harpalus retractus LeConte, 1863


 * Subgenus Pseudoophonus Motschulsky, 1844


 * Harpalus actiosus Casey, 1914
 * Harpalus aenigma (Tschitscherine, 1897)
 * Harpalus alienus Bates, 1878
 * Harpalus aogashimensis (Habu, 1957)
 * Harpalus azumai Habu, 1968
 * Harpalus babai Habu, 1973
 * Harpalus calceatus (Duftschmid, 1812)
 * Harpalus caliginosus (Fabricius, 1775)
 * Harpalus capito A.Morawitz, 1862
 * Harpalus compar LeConte, 1847
 * Harpalus coreanus (Tschitscherine, 1895)
 * Harpalus davidi (Tschitscherine, 1897)
 * Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996
 * Harpalus eous Tschitscherine, 1901
 * Harpalus erythropus Dejean, 1829
 * Harpalus faunus Say, 1823
 * Harpalus fokienensis Schauberger, 1930
 * Harpalus griseus (Panzer, 1796)
 * Harpalus hatchi Ball & Anderson, 1962
 * Harpalus hauserianus Schauberger, 1929
 * Harpalus indicus Bates, 1891
 * Harpalus jureceki (Jedlicka, 1928)
 * Harpalus katiae F.Battoni, 1985
 * Harpalus liobasis Chaudoir, 1868
 * Harpalus meghalayensis Kataev, 2001
 * Harpalus meridianus Andrewes, 1923
 * Harpalus paratus Casey, 1924
 * Harpalus pastor Motschulsky, 1844
 * Harpalus pensylvanicus (DeGeer, 1774)
 * Harpalus poncei Will, 2002
 * Harpalus protractus Casey, 1914
 * Harpalus pseudohauserianus Kataev, 2001
 * Harpalus pseudophonoides Schauberger, 1930
 * Harpalus roninus Bates, 1873
 * Harpalus rufipes (DeGeer, 1774)
 * Harpalus sericatus Tschitscherine, 1906
 * Harpalus simplicidens Schauberger, 1929
 * Harpalus singularis Tschitscherine, 1906
 * Harpalus sinicus Hope, 1845
 * Harpalus suensoni Kataev, 1997
 * Harpalus texanus Casey, 1914
 * Harpalus tridens A.Morawitz, 1862
 * Harpalus ussuriensis Chaudoir, 1863
 * Harpalus vagans LeConte, 1865
 * Harpalus yinchuanensis Huang, 1993


 * Subgenus Semiophonus Schauberger, 1933
 * Harpalus signaticornis (Duftschmid, 1812)


 * Subgenus Zangoharpalus Huang, 1998
 * Harpalus mariae Kataev, 1997
 * Harpalus microdemas Schauberger, 1932
 * Harpalus praticola Bates, 1891
 * Harpalus pseudotinctulus Schauberger, 1932
 * Harpalus tinctulus Bates, 1873


 * Extinct, not assigned to subgenus


 * †Harpalus abolitus C. & L.Heyden, 1866
 * †Harpalus anactus Giebel, 1856
 * †Harpalus bruckmanni Heer, 1862
 * †Harpalus burmeisteri Giebel, 1856
 * †Harpalus conditus Scudder, 1900
 * †Harpalus constrictus Heer, 1862
 * †Harpalus deletus Oustalet, 1864
 * †Harpalus dilatatus Theobald, 1937
 * †Harpalus ewaldi Giebel, 1856
 * †Harpalus excavatus Forster, 1891
 * †Harpalus heeri Giebel, 1856
 * †Harpalus knorri Giebel, 1856
 * †Harpalus lheritieri Piton, 1940
 * †Harpalus liasinus Giebel, 1856
 * †Harpalus maceratus Wickham, 1911
 * †Harpalus nero Oustalet, 1864
 * †Harpalus nitens Birket-Smith, 1977
 * †Harpalus nuperus Scudder, 1900
 * †Harpalus obtusus Forster, 1891
 * †Harpalus oustaleti Meunier, 1915
 * †Harpalus pleistocenicus Lomnicki, 1894
 * †Harpalus redivivus Wickham, 1917
 * †Harpalus schlotheimi Giebel, 1856
 * †Harpalus sinis Heer, 1847
 * †Harpalus stierlini Heer, 1862
 * †Harpalus stygius Heer, 1862
 * †Harpalus tabidus Heer, 1847
 * †Harpalus tardigradus Heer, 1862
 * †Harpalus ulomaeformis Wickham, 1917
 * †Harpalus veterum Cockerell, 1920
 * †Harpalus whitfieldi Scudder, 1900