Holepyris

Holepyris is a genus of hymenopteran parasitoids in the family Bethylidae. These species use small beetle larvae as hosts for their larvae. Prior to a 2022 study that found this genus and other genera within the subfamily Epyrinae to be polyphyletic, there were 148 species in the genus Holepyris. Following a reclassification aiming at creating monophyletic genera of Epyrinae, 131 Holepyris species were placed in the genus Rysepyris (including Holepyris sylvanidis, now Rysepyris sylvanidis, a species studied as a biocontrol agent of some storage pests), and three Formosiepyris species were placed in the genus Holepyris. As a result, there are now 23 species in the genus Holepyris:

Species

 * Holepyris africanus Kieffer, 19041
 * Holepyris agraensis (Kurian, 1955)
 * Holepyris algoaensis Kieffer, 1911
 * Holepyris angusticollis Kieffer, 1906
 * Holepyris bergevini Duchaussoy, 1916[1914]
 * Holepyris bidentatus Kieffer, 1904
 * Holepyris crenulatus Kieffer, 1906
 * Holepyris fasciipennis Kieffer, 1906
 * Holepyris fuscipennis Kieffer, 1906
 * Holepyris levicollis Kieffer, 1906
 * Holepyris lineatus Kieffer, 1906
 * Holepyris maculipennis (Marshall, 1874)
 * Holepyris marishi (Terayama, 2004)
 * Holepyris natalensis Kieffer, 1913
 * Holepyris orientalis Kieffer, 1906
 * Holepyris punctaticollis Kieffer, 1906
 * Holepyris rufitarsis Kieffer, 1906
 * Holepyris rugulosus (Xu & He, 2005)
 * Holepyris shiva (Terayama, 2004)
 * Holepyris takasago (Terayama, 2004)
 * Holepyris tibialis Kieffer, 1906
 * Holepyris vietnamensis (Tsujii et al., 2015)
 * Holepyris vittatus Turner, 1928