Holothyrida

The Holothyrida are a small order of mites in the superorder Parasitiformes. No fossils are known. With body lengths of more than 2 mm they are relatively large mites, with a heavily sclerotized body. It is divided into three families, Allothyridae, Holothyridae, and Neothyridae. In a 1998 experimental study, members of the family Allothyridae were found to ignore living animals but readily fed on the body fluids of dead arthropods, making them scavengers.

The order has a distribution largely confined to former Gondwanan landmasses. They are the sister group to Ixodida (ticks).

Allothyridae
Allothyridae van der Hammen, 1972 — Australia, New Zealand


 * Allothyrus van der Hammen, 1961
 * Allothyrus australasiae (Womersley, 1935)
 * Allothyrus constrictus (Domrow, 1955)


 * Australothyrus van der Hammen, 1983
 * Australothyrus ocellatus van der Hammen, 1983

Holothyridae
Holothyridae Thorell, 1882 Sri Lanka, Indian Ocean islands, New Guinea, New Caledonia


 * Sternothyrus Lehtinen, 1995
 * Sternothyrus braueri (Thon, 1905) — Seychelles


 * Lindothyrus Lehtinen, 1995
 * Lindothyrus elongatus Lehtinen, 1995 — Lord Howe Island
 * Lindothyrus rubellus Lehtinen, 1995 — New Caledonia


 * Indothyrus Lehtinen, 1995
 * Indothyrus greeni Lehtinen, 1995 — Sri Lanka


 * Haplothyrus Lehtinen, 1995
 * Haplothyrus expolitissimus (Berlese, 1924) — New Caledonia
 * Haplothyrus hyatti Lehtinen, 1995 — unknown locality


 * Holothyrus Gervais, 1842 — Mauritius
 * Holothyrus coccinella Gervais, 1842
 * Holothyrus legendrei Hammen, 1983


 * Hammenius Lehtinen, 1981
 * Hammenius armatus (Canestrini, 1897) — Tamara Island (Aitape): New Guinea
 * Hammenius berlesei (Lehtinen, 1995) — New Guinea
 * Hammenius braueri (Thon, 1906)
 * Hammenius fujuge Lehtinen, 1981 — New Guinea (Central District, Oro Province)
 * Hammenius grandjeani (Hammen, 1961) — Mount Bosavi: New Guinea
 * Hammenius holthuisi van der Hammen, 1983
 * Hammenius ingii Lehtinen, 1981 — New Guinea
 * Hammenius insularis Lehtinen, 1995 — Louisiade Archipelago: New Guinea
 * Hammenius longipes (Thorell, 1882) — Fly River, New Guinea (?)
 * Hammenius mendi (Lehtinen, 1995) — Strickland River: New Guinea
 * Hammenius montanus Hammen, 1983 — Irian Jaya
 * Hammenius niger (Thon, 1906)

Neothyridae
Neothyridae Lehtinen, 1981 Northern South America and the Caribbean


 * Diplothyrus Lehtinen, 1999 Brazil, French Guiana
 * Diplothyrus schubarti Lehtinen, 1999
 * Diplothyrus lecorrei Klompen 2010
 * Diplothyrus lehtineni Vázquez & de Araújo & Feres 2016
 * Neothyrus Lehtinen, 1981 Peru
 * Neothyrus ana Lehtinen, 1981
 * Caribothyrus Kontschán & Mahunka 2004 Dominican Republic
 * Caribothyrus barbatus Kontschán & Mahunka 2004