Milnesium

Milnesium is a genus of tardigrades. It is rather common, being found in a wide variety of habitats across the world. It has a fossil record extending back to the Cretaceous, the oldest species found so far (M. swolenskyi) is known from Turonian stage deposits on the east coast of the United States. Milnesiums are one of the most desiccation and radiation-resistant invertebrates on Earth because of their unique ability to transform into a "tun" state and utilize intrinsically disordered proteins when experiencing extreme environments.

Species
Described species include:


 * Milnesium alabamae Wallendorf & Miller, 2009
 * Milnesium almatyense Tumanov, 2006
 * Milnesium alpigenum Ehrenberg, 1853
 * Milnesium antarcticum Tumanov, 2006
 * Milnesium argentinum Roszkowska, Ostrowska & Kaczmarek, 2015
 * Milnesium asiaticum Tumanov, 2006
 * Milnesium barbadosense Meyer & Hinton, 2012
 * Milnesium beasleyi Kaczmarek, Jakubowska & Michalczyk, 2012
 * Milnesium beatae Roszkowska, Ostrowska & Kaczmarek, 2015
 * Milnesium berladnicorum Ciobanu, Zawierucha, Moglan & Kaczmarek, 2014
 * Milnesium bohleberi Bartels, Nelson, Kaczmarek & Michalczyk, 2014
 * Milnesium brachyungue Binda & Pilato, 1990
 * Milnesium cassandrae Moreno-Talamantes, Roszkowska, García-Aranda, Flores-Maldonado & Kaczmarek, 2019
 * Milnesium burgessi Schlabach, Donaldson, Hobelman, Miller & Lowman, 2018
 * Milnesium dornensis Ciobanu, Roszkowska & Kaczmarek, 2015
 * Milnesium dujiangensis Yang, 2003
 * Milnesium eurystomum Maucci, 1991
 * Milnesium fridae Moreno-Talamantes, León-Espinosa, García-Aranda, Flores-Maldonado & Kaczmarek, 2020
 * Milnesium granulatum Ramazzotti, 1962
 * Milnesium jacobi Meyer & Hinton, 2010
 * Milnesium katarzynae Kaczmarek, Michalczyk & Beasley, 2004
 * Milnesium kogui Londoño, Daza, Caicedo, Quiroga & Kaczmarek, 2015
 * Milnesium krzysztofi Kaczmarek & Michalczyk, 2007
 * Milnesium lagniappe Meyer, Hinton & Dupré, 2013
 * Milnesium longiungue Tumanov, 2006
 * Milnesium minutum Pilato & Lisi, 2016
 * Milnesium quadrifidum Nederström, 1919
 * Milnesium rastrum Suzuki, Sugiura, Tsujimoto, and McInnes, 2023
 * Milnesium reductum Tumanov, 2006
 * Milnesium reticulatum Pilato, Binda & Lisi, 2002
 * Milnesium sandrae Pilato & Lisi, 2016
 * Milnesium shilohae Meyer, 2015
 * Milnesium swansoni Young, Chappell, Miller & Lowman, 2016
 * † Milnesium swolenskyi Bertolani & Grimaldi, 2000
 * Milnesium tardigradum Doyère, 1840
 * Milnesium tetralamellatum Pilato & Binda, 1991
 * Milnesium tumanovi Pilato, Sabella & Lisi, 2016
 * Milnesium validum Pilato, Sabella, D'Urso & Lisi, 2017
 * Milnesium variefidum Morek, Gąsiorek, Stec, Blagden & Michalczyk, 2016
 * Milnesium vorax Pilato, Sabella & Lisi, 2016
 * Milnesium zsalakoae Meyer & Hinton, 2010