Pentatominae

Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 4937 species classified in 938 genera. Species in this subfamily are phytophages and several of them are considered agricultural pests. Some invasive pentatomines such as Halyomorpha halys and Bagrada hilaris have been considered household pests. Higher systematics of the group have been revised by Rider et al.

Tribes and genera
Tribes within this subfamily include:


 * Aelini Douglas & Scott, 1865
 * 1) Aelia Fabricius, 1803
 * 2) Aeliopsis Bergevin, 1931
 * 3) Neottiglossa Kirby, 1837
 * Aeptini Stål, 1871
 * 1) Aeliosoma Baehr, 1985
 * 2) Aeptus Dallas, 1851
 * 3) Caridillus Bergroth, 1909
 * 4) Eribotes (bug) Stål, 1867
 * 5) Hillieria Distant, 1910
 * 6) Mariomella Faúndez & Rider, 2019
 * 7) Menestheus (bug) Stål, 1867
 * 8) Paramenestheus Breddin, 1900
 * 9) Pseudaelia Distant, 1910
 * Aeschrocorini Distant, 1902
 * 1) Aeschrocoris Bergroth, 1887
 * 2) Aeschrus Spinola, 1850
 * 3) Chraesus Cachan, 1952
 * 4) Geomorpha (bug) Bergroth, 1893
 * 5) Risbecella Schouteden, 1942
 * 6) Scylax (bug) Distant, 1887
 * 7) Tyoma Spinola, 1850
 * 8) Tyomana Miller, 1952
 * Agaeini Cachan, 1952
 * 1) Agaeus Dallas, 1851
 * Agonoscelidini Atkinson, 1888
 * 1) Agonoscelis Spinola, 1837
 * Amyntorini Distant, 1902
 * 1) Ambohicorypha Cachan, 1952
 * 2) Belopis Distant, 1879
 * 3) Bolaca Walker, 1867
 * 4) Halyabbas Distant, 1900
 * 5) Madecorypha Cachan, 1952
 * 6) Sennertus Distant, 1900
 * Antestiini Distant, 1902 - selected genera:
 * Antestia Stål, 1865
 * Antestiopsis Leston, 1952
 * Dryadocoris Kirkaldy, 1909
 * Plautia Stål, 1864
 * Aulacetrini Mulsant & Rey, 1866
 * 1) Holcogaster Fieber, 1860
 * Axiagastini Atkinson, 1888
 * 1) Acesines Stål, 1876
 * 2) Axiagastus Dallas, 1851
 * 3) Indrapura (bug) China, 1928
 * 4) Oncotropis Stål, 1871
 * Bathycoeliini Atkinson, 1888
 * 1) Bathycoelia Amyot & Serville, 1843
 * Cappaeini Atkinson, 1888
 * 1) Adelolcus Bergroth, 1893
 * 2) Benia (bug) Schouteden, 1916
 * 3) Boerias Kirkaldy, 1909
 * 4) Cappaea Ellenrieder, 1862
 * 5) Caura (bug) Stål, 1865
 * 6) Cauromorpha Jeannel, 1913
 * 7) Erlangerella Schouteden, 1905
 * 8) Halycorypha Jeannel, 1913
 * 9) Halydicoris Jeannel, 1913
 * 10) Halyoides Cachan, 1952
 * 11) Halyomorpha Mayr, 1864
 * 12) Homalogonia Jakovlev, 1876
 * 13) Hymenomaga Karsch, 1892
 * 14) Lerida (bug) Karsch, 1894
 * 15) Leridella Jeannel, 1913
 * 16) Lokaia Linnavuori, 1975
 * 17) Mabusana Distant, 1912
 * 18) Massocephalus Dallas, 1851
 * 19) Paralerida Linnavuori, 1982
 * 20) Prytanicoris Gross, 1978
 * 21) Tolumnia Stål, 1867
 * 22) Tripanda Berg, 1899
 * 23) Tropicorypha Mayr, 1864
 * 24) Veterna Stål, 1865
 * Carpocorini Mulsant & Rey, 1866 selected genera:
 * Antheminia Mulsant & Rey, 1866
 * Carpocoris Kolenati, 1846
 * Chlorochroa Stål, 1872
 * Dolycoris Mulsant & Rey, 1866
 * Graphorn Faúndez, Rider, & Carvajal, 2017
 * Peribalus Mulsant & Rey, 1866
 * Rubiconia Dohrn, 1860
 * Thestral Faúndez & Rider, 2014
 * Catacanthini Atkinson, 1888
 * 1) Arocera Spinola, 1837
 * 2) Boea (bug) Walker, 1867
 * 3) Catacanthus Spinola, 1837
 * 4) Chalcocoris Dallas, 1851
 * 5) Rhyssocephala Rider, 1992
 * 6) Runibia Stål, 1861
 * 7) Vitruvius (bug) Distant, 1901
 * 8) Vulsirea Spinola, 1837
 * Caystrini Ahmad & Afzal, 1979
 * 1) Caystrus Stål, 1861
 * 2) Cephaloplatus White, 1842
 * 3) Heissocoris Ahmad & Afzal, 1979
 * 4) Hippotiscus Bergroth, 1906
 * 5) Kyrtalus Van Duzee, 1929
 * 6) Lyalocoris Zahid, 2006
 * 7) Mcdonaldiana Zahid, 2006
 * 8) Neodius (Stål, 1867)
 * 9) Neoparodius Zahid, 2006
 * 10) Neoriazocoris Zahid, 2006
 * 11) Paramecocoris Stål, 1853
 * 12) Parodius (bug) Distant, 1900
 * 13) Praetextatus (bug) Distant, 1901
 * 14) Riazocoris Ahmad & Afzal, 1979
 * 15) Valescus Distant, 1901
 * Chlorocorini Rider, Greve, Schwertner & Grazia, 2018
 * 1) Arvelius Spinola, 1837
 * 2) Chlorocoris Spinola, 1837
 * 3) Chloropepla Stål, 1867
 * 4) Eludocoris Thomas, 1992
 * 5) Fecelia Stål, 1872
 * 6) Mayrinia Horváth, 1925
 * 7) Rhyncholepta Bergroth, 1911
 * Coquereliini Cachan, 1952
 * 1) Cleoqueria Cachan, 1952
 * 2) Coquerelia Signoret, 1861
 * 3) Coquerelidea Reuter, 1887
 * 4) Neocoquerelidea Cachan, 1952
 * Degonetini Azim & Shafee, 1984
 * 1) Degonetus Distant, 1902
 * Diemeniini Kirkaldy, 1909 - selected genera:
 * Commius (bug) Stål, 1876
 * Diemenia (bug) Spinola, 1850
 * Diplostirini Distant, 1902
 * 1) Ambiorix (bug) Stål, 1876
 * 2) Diplostira Dallas, 1851
 * Diploxyini Atkinson, 1888
 * 1) Acoloba Spinola, 1850
 * 2) Aeladria Cachan, 1952
 * 3) Diploxys Amyot & Serville, 1843
 * Eurysaspidini Atkinson, 1888
 * 1) Eurysaspis Signoret, 1851
 * 2) Flaminia (bug) Stål, 1865
 * Eysarcorini Mulsant & Rey, 1866
 * 1) Adria (bug) Stål, 1876
 * 2) Aspavia Stål, 1865
 * 3) Cachanocoris Rider, 1998
 * 4) Carbula Stål, 1864
 * 5) Corisseura Cachan, 1952
 * 6) Cratonotus Distant, 1879
 * 7) Durmia Stål, 1865
 * 8) Dymantiscus Hsiao, 1981
 * 9) Empiesta Linnavuori, 1982
 * 10) Eysarcoris Hahn, 1834
 * 11) Hermolaus Distant, 1902
 * 12) Mulungua Linnavuori, 1982
 * 13) Pseudolerida Schouteden, 1958
 * 14) Sepontia Stål, 1865
 * 15) Sepontiella Miyamoto, 1990
 * 16) Spermatodes Bergroth, 1914
 * 17) Stagonomus Gorski, 1852
 * Halyini Amyot & Serville, 1843 - selected genera:
 * Brochymena Amyot & Serville, 1843
 * Hoplistoderini Atkinson, 1888
 * 1) Alcimocoris Bergroth, 1891
 * 2) Glottaspis Bergroth, 1918
 * 3) Hoplistodera Westwood, 1837
 * 4) Paracritheus Bergroth, 1891
 * 5) Paralcimocoris Zhang & Lin, 1988
 * 6) Stachyomia Stål, 1871
 * 7) Taurodes Dallas, 1851
 * Lestonocorini Ahmad & Mohammad, 1980
 * 1) Dardjilingia Yang, 1936
 * 2) Gynenica Dallas, 1851
 * 3) Lestonocoris Ahmad & Mohammad, 1980
 * 4) Neogynenica Yang, 1935
 * 5) Umgababa (bug) Leston, 1953
 * Mecideini Distant, 1902
 * 1) Mecidea Dallas, 1851
 * Memmiini Cachan, 1952
 * 1) Memmia Stål, 1865
 * Menidini Atkinson, 1888
 * 1) Actuarius (bug) Distant, 1901
 * 2) Aegaleus (bug) Stål, 1876
 * 3) Amphimachus (bug) Stål, 1867
 * 4) Anchesmus Stål, 1876
 * 5) Apines Dallas, 1851
 * 6) Aspideurus Signoret, 1880
 * 7) Banya (bug) Schouteden, 1916
 * 8) Brachycoris Stål, 1871
 * 9) Carvalhocoris Leston, 1953
 * 10) Cresphontes (bug) Stål, 1867
 * 11) Dabessus Distant, 1902
 * 12) Decellella Schouteden, 1964
 * 13) Desertomenida Kiritshenko, 1914
 * 14) Dunnius Distant, 1902
 * 15) Elanela Rolston, 1980
 * 16) Eurymenida Bergroth, 1917
 * 17) Gwea Schouteden, 1958
 * 18) Keleacoris Rider & Rolston, 1995
 * 19) Lathraedoeus Breddin, 1900
 * 20) Menida Motschulsky, 1861
 * 21) Neostrachia Saunders, 1877
 * 22) Phavorinus Distant, 1902
 * 23) Rio (bug) Kirkaldy, 1909
 * 24) Saceseurus Breddin, 1900
 * 25) Sciomenida Gross, 1976
 * 26) Solenocoris Oshanin, 1922
 * 27) Udonga Distant, 1921
 * Myrocheini Stål, 1871
 * 1) Aednulus Breddin, 1901
 * 2) Aednus Dallas, 1851
 * 3) Arniscus Distant, 1899
 * 4) Delegorguella Spinola, 1850
 * 5) Dictyotus Dallas, 1851
 * 6) Dictytotus Dallas, 1851
 * 7) Dissocolpus Bergroth, 1906
 * 8) Dollingiana Ahmad & Kamaluddin, 1986
 * 9) Dymantis Stål, 1861
 * 10) Ennius (bug) Stål, 1861
 * 11) Erachtheus Stål, 1861
 * 12) Humria Linnavuori, 1975
 * 13) Jitka (bug) Kment, 2015
 * 14) Laprius Stål, 1861
 * 15) Lodosia Ahmad & Önder, 1996
 * 16) Munshiana Ahmad & Kamaluddin, 1987
 * 17) Myrochea Amyot & Serville, 1843
 * 18) Neococalus Bergroth, 1891
 * 19) Neodorpius Ahmad & Afzal, 1989
 * 20) Paradictyotus Gross, 1975
 * 21) Pretorius (bug) Distant, 1898
 * 22) Stysicoris Ahmad & Kamaluddin, 1985
 * 23) Tholosanus Distant, 1899
 * 24) Utheria Gross, 1975
 * Nealeriini Cachan, 1952
 * 1) Nealeria Bergroth, 1893
 * 2) Paraleria Reuter, 1887
 * Nezarini Atkinson, 1888 - selected genera:
 * Nezara Amyot et Serville, 1843
 * Palomena Mulsant et Rey, 1866
 * Opsitomini Cachan, 1952
 * 1) Opsitoma Cachan, 1952
 * Pentamyrmexini Rider & Brailovsky, 2014 (monotypic, Thailand)
 * Pentamyrmex Rider & Brailovsky, 2014
 * Pentatomini Leach, 1815 - selected genera:
 * Pentatoma Olivier, 1789
 * Phricodini Cachan, 1952
 * 1) Phricodus Spinola, 1839
 * Piezodorini Atkinson, 1888
 * 1) Anaximenes (bug) Stål, 1876
 * 2) Chaubattiana Distant, 1912
 * 3) Pausias (bug) Jakovlev, 1905
 * 4) Piezodorus Fieber, 1860
 * Procleticini Pennington, 1920
 * 1) Aleixus McDonald, 1981
 * 2) Brepholoxa Van Duzee, 1904
 * 3) Dendrocoris Bergroth, 1891
 * 4) Lobepomis Berg, 1891
 * 5) Neoderoploa Pennington, 1922
 * 6) Odmalea Bergroth, 1914
 * 7) Parodmalea Rider, 1994
 * 8) Procleticus Berg, 1891
 * 9) Pseudobebaeus Distant, 1911
 * 10) Terania Pirán, 1963
 * 11) Thoreyella Spinola, 1850
 * Rhynchocorini Stål, 1871
 * 1) Araducta Walker, 1867
 * 2) Avicenna (bug) Distant, 1901
 * 3) Biprorulus Breddin, 1900
 * 4) Cuspicona Dallas, 1851
 * 5) Diaphyta Bergroth, 1891
 * 6) Everardia (bug) Gross, 1975
 * 7) Hoffmanseggiella Spinola, 1850
 * 8) Koogobatha Distant, 1910
 * 9) Morna (bug) Stål, 1867
 * 10) Mygoodano Distant, 1910
 * 11) Ocirrhoe Stål, 1876
 * 12) Parocirrhoe Gross, 1975
 * 13) Pegala Stål, 1867
 * 14) Petalaspis Bergroth, 1916
 * 15) Pugione Stål, 1867
 * 16) Rhynchocoris Westwood, 1837
 * 17) Utana Distant, 1900
 * 18) Vitellus Stål, 1865
 * Rolstoniellini'' Rider, 1997
 * 1) Agathocles (bug) Stål, 1876
 * 2) Amasenus Stål, 1863
 * 3) Amatembuna Distant, 1910
 * 4) Critheus Stål, 1867
 * 5) Exithemus Distant, 1902
 * 6) Rolstoniellus Rider, 1997
 * Sciocorini Amyot & Serville, 1843 selected genera:
 * Dyroderes Spinola, 1837
 * Sciocoris Fallén, 1829
 * Strachiini Mulsant & Rey, 1866 selected genera:
 * Afrania Stål, 1865
 * Bagrada Stål, 1862
 * Eurydema Laporte de Castelnau, 1833
 * Murgantia (bug) Stål, 1862
 * Stenozygum Fieber, 1861
 * Trochiscocoris Reuter, 1890
 * Triplatyxini Cachan, 1952
 * 1) Anoano Cachan, 1952
 * 2) Nene (bug) Kment, 2015
 * 3) Tricompastes Cachan, 1952
 * 4) Triplatyx Horváth, 1904

Genera incertae sedis

 * 1) Amphidexius Bergroth, 1918
 * 2) Bachesua Gross, 1975
 * 3) Birna McDonald, 2006
 * 4) Brizocoris Gross, 1975
 * 5) Buthumka Gross, 1975
 * 6) Cooperocoris Gross, 1975
 * 7) Dysnoetus Bergroth, 1918
 * 8) Eremophilacoris Gross, 1975
 * 9) Gadarscama Reuter, 1887
 * 10) Ippatha Distant, 1910
 * 11) Kitsonia Gross, 1975
 * 12) Kitsoniocoris Rider, 1998
 * 13) Kumbutha Distant, 1910
 * 14) Lakhonia Yang, 1936
 * 15) Macrocarenoides Gross, 1975
 * 16) Macrocarenus Stål, 1873
 * 17) Mcphersonarcys Thomas, 2012
 * 18) Muscanda Walker, 1868
 * 19) Ochisme Kirkaldy, 1904
 * 20) Ooldeon Gross, 1975
 * 21) Patanius Rolston, 1987
 * 22) Pirricoris Gross, 1975
 * 23) Poecilotoma Dallas, 1851
 * 24) Riaziana Hasan, Afzal & Ahmad, 1989
 * 25) Senectius Rolston, 1987
 * 26) Tepperocoris Gross, 1975
 * 27) Thryptomenecoris Gross, 1975
 * 28) Trachyops Dallas, 1851
 * 29) Turrubulana Distant, 1910
 * 30) Xiengia Distant, 1921
 * 31) Zouicoris Zheng, 1986

Notable species

 * Acrostemum hilare – green stink bug
 * Bagrada hilaris – bagrada bug
 * Bathycoelia thalassina - African cocoa shield bug
 * Halyomorpha halys – brown marmorated stink bug
 * Nezara viridula - green vegetable bug
 * Pentatoma rufipes – forest bug
 * Piezodorus lituratus – gorse shield bug