Phylinae

Phylinae is a subfamily of the plant bug family Miridae. Species of this family are found worldwide.

In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, and the tribe Pronotocrepini a synonym of the subtribe Cremnorrhina. New tribes were created under the new or resurrected names Decomiini, Exocarpocorini, Cremnorrhini, Nasocorini, and Semiini. The revised subfamily consists of nine tribes containing around 400 genera and over 2000 species.


 * Subfamily Phylinae Douglas & Scott, 1865
 * Tribe Cremnorrhinini Reuter, 1883 (sometimes spelled "Cremnorrhini")
 * Subtribe Coatonocapsina Schuh & Menard, 2013
 * Subtribe Cremnorrhina Reuter, 1883
 * Tribe Decomiini Schuh & Menard, 2013
 * Tribe Exaeretini Futon, 1875
 * Tribe Hallodapini Van Duzee, 1916
 * Tribe Leucophoropterini Schuh, 1974
 * Subtribe Leucophoropterina Schuh, 1974
 * Subtribe Tuxedoina Schuh & Menard, 2013
 * Tribe Nasocorini Reuter, 1883
 * Tribe Phylini Douglas & Scott, 1865
 * Subtribe Keltoniina Schuh & Menard, 2013
 * Subtribe Oncotylina Douglas & Scott, 1865
 * Subtribe Phylina Douglas & Scott, 1865
 * Tribe Pilophorini Douglas & Scott, 1876
 * Tribe Semiini Knight, 1923
 * Subtribe Exocarpocorina Schuh & Menard, 2013
 * Subtribe Semiina Knight, 1923

Genera
Phylinae contains the following genera:


 * Subfamily Phylinae Douglas & Scott, 1865
 * Tribe Cremnorrhinini Reuter, 1883
 * Subtribe Coatonocapsina Schuh & Menard, 2013
 * Genus Austropsallus Schuh, 1974
 * Genus Capecapsus Schuh, 1974
 * Genus Coatonocapsus Schuh, 1974
 * Genus Heterocapillus Wagner, 1960
 * Genus Parasciodema Poppius, 1914
 * Subtribe Cremnorrhinina Reuter, 1883
 * Genus Amblytylus Fieber, 1858
 * Genus Brachyceratocoris Knight, 1968
 * Genus Calidroides Schwartz, 2005
 * Genus Coquillettia Uhler, 1890
 * Genus Cremnorrhinus Reuter, 1880
 * Genus Dacota Uhler, 1872
 * Genus Denticulophallus Schuh, 1974
 * Genus Ethelastia Reuter, 1876
 * Genus Euderon Puton, 1888
 * Genus Excentricoris Carvalho, 1955
 * Genus Guentherocoris Schuh & Schwartz, 2004
 * Genus Harpocera Curtis, 1838
 * Genus Leutiola Wyniger, 2012
 * Genus Lopidodenus V. Putshkov, 1974
 * Genus Lopus Hahn, 1833
 * Genus Macrotylus Fieber, 1858
 * Genus Orectoderus Uhler, 1876
 * Genus Pachyxyphus Fieber, 1858
 * Genus Paralopus Wagner, 1957
 * Genus Pronotocrepis Knight, 1929
 * Genus Shendina Linnavuori, 1975
 * Genus Strophopoda Van Duzee, 1916
 * Genus Teleorhinus Uhler, 1890
 * Genus Ticua Wyniger, 2012
 * Genus Utopnia Reuter, 1881
 * Genus Zinjolopus Linnavuori, 1975
 * Tribe Decomiini Schuh & Menard, 2013
 * Genus Aurantiocoris Schuh & Schwartz, 2004
 * Genus Decomia Poppius, 1915
 * Genus Decomioides Schuh, 1984
 * Genus Malaysiamiris Schuh, 1984
 * Genus Malaysiamiroides Schuh, 1984
 * Genus Rubrocuneocoris Schuh, 1984
 * Tribe Exaeretini Futon, 1875
 * Genus Anonychiella Reuter, 1912
 * Genus Aphaenophyes Reuter, 1899
 * Genus Atractotomimus Kiritshenko, 1952
 * Genus Auchenocrepis Fieber, 1858
 * Genus Brendaphylus Yasunaga, 2013
 * Genus Camptotylus Fieber, 1860
 * Genus Camptozorus Kerzhner, 1996
 * Genus Chrysochnoodes Reuter, 1901
 * Genus Compsonannus Reuter, 1902
 * Genus Eumecotarsus Kerzhner, 1962
 * Genus Eurycranella Reuter, 1904
 * Genus Frotaphylus Carvalho, 1984
 * Genus Gonoporomiris Henry & Schuh, 2002
 * Genus Hadrophyes Puton, 1874
 * Genus Hyalopsallus Carvalho & Schaffner, 1973
 * Genus Megalodactylus Fieber, 1858
 * Genus Moissonia Reuter, 1894
 * Genus Opuna Kirkaldy, 1902
 * Genus Pastocoris Reuter, 1879
 * Genus Psallopsis Reuter, 1901
 * Genus Randallopsallus Yasunaga, 2013
 * Genus Tuponia Reuter, 1875
 * Genus Voruchiella Poppius, 1912
 * Genus Yotvata Linnavuori, 1964
 * Tribe Hallodapini Van Duzee, 1916
 * Genus Acrorrhinium Noualhier, 1895
 * Genus Aeolocoris Reuter, 1903
 * Genus Alloeomimus Reuter, 1910
 * Genus Artchawakomius Yasunaga, 2012
 * Genus Aspidacanthus Reuter, 1901
 * Genus Auricillocoris Schuh, 1984
 * Genus Azizus Distant, 1910
 * Genus Bibundiella Poppius, 1914
 * Genus Boopidella Reuter, 1907
 * Genus Carinogulus Schuh, 1974
 * Genus Chaetocapsus Poppius, 1914
 * Genus Clapmarius Distant, 1904
 * Genus Cleotomiris Schuh, 1984
 * Genus Cleotomiroides Schuh, 1984
 * Genus Cremnocephalus Eieber, 1860
 * Genus Cyrtopeltocoris Reuter, 1876
 * Genus Diocoris Kirkaldy, 1902
 * Genus Eminoculus Schuh, 1974
 * Genus Formicopsella Poppius, 1914
 * Genus Gampsodema Odhiambo, 1960
 * Genus Glaphyrocoris Reuter, 1903
 * Genus Hadrodapus Linnavuori, 1996
 * Genus Hallodapomimus Herczek, 2000
 * Genus Hallodapus Eieber, 1858
 * Genus Ifephylus Linnavuori, 1993
 * Genus Kapoetius Schmitz, 1969
 * Genus Laemocoris Reuter, 1879
 * Genus Leaina Linnavuori, 1974
 * Genus Leptomimus Herczek & Popov, 2010
 * Genus Lestonisca Carvalho, 1988
 * Genus Linacoris Carvalho, 1983
 * Genus Lissocapsus Bergroth, 1903
 * Genus Malgacheocoris Carvalho, 1952
 * Genus Mimocoris J. Scott, 1872
 * Genus Myombea China & Carvalho, 1951
 * Genus Myrmicomimus Reuter, 1881
 * Genus Neolaemocoris Wagner, 1975
 * Genus Omphalonotus Reuter, 1876
 * Genus Pangania Poppius, 1914
 * Genus Paralaemocoris Linnavuori, 1964
 * Genus Phoradendrepulus Polhemus & Polhemus, 1985
 * Genus Podullahas Schuh, 1984
 * Genus Pongocoris Linnavuori, 1975
 * Genus Ribautocapsus Wagner, 1962
 * Genus Ruwaba Linnavuori, 1975
 * Genus Skukuza Schuh, 1974
 * Genus Sohenus Distant, 1910
 * Genus Syngonus Bergroth, 1926
 * Genus Systellonotidea Poppius, 1914
 * Genus Systellonotopsis Poppius, 1914
 * Genus Systellonotus Pieber, 1858
 * Genus Trichophorella Reuter, 1905
 * Genus Trichophthalmocapsus Poppius, 1914
 * Genus Vitsikamiris Polhemus, 1994
 * Genus Wygomiris Schuh, 1984
 * Genus Zaratus Distant, 1909
 * Tribe Leucophoropterini Schuh, 1974
 * Subtribe Leucophoropterina Schuh, 1974
 * Genus Abuyogocoris Schuh, 1984
 * Genus Aitkenia Carvalho & Gross, 1982
 * Genus Arafuramiris Schuh, 1984
 * Genus Ausejanus Menard & Schuh, 2011
 * Genus Austrodapus Menard & Schuh, 2011
 * Genus Biromiris Schuh, 1984
 * Genus Blesingia Carvalho & Gross, 1982
 * Genus Collessicoris Carvalho & Gross, 1982
 * Genus Ctypomiris Schuh, 1984
 * Genus Gulacapsus Schuh, 1984
 * Genus Johnstonsonius Menard & Schuh, 2011
 * Genus Leucophoroptera Poppius, 1921
 * Genus Missanos Menard & Schuh, 2011
 * Genus Neaitkenia Menard & Schuh, 2011
 * Genus Neoleucophoroptera Menard & Schuh, 2011
 * Genus Papuamimus Schuh, 1984
 * Genus Papuamiroides Menard & Schuh, 2011
 * Genus Pseudohallodapocoris Schuh, 1984
 * Genus Sejanus Distant, 1910
 * Genus Solomonomimus Schuh, 1984
 * Genus Transleucophoroptera Menard & Schuh, 2011
 * Genus Trichocephalocapsus Schuh, 1984
 * Genus Waterhouseana Carvalho, 1973
 * Subtribe Tuxedoina Schuh & Menard, 2013
 * Genus Ephippiocoris Poppius, 1912
 * Genus Pseudophylus Yasunaga, 1999
 * Genus Tuxedo Schuh, 2001
 * Tribe Nasocorini Reuter, 1883
 * Genus Adenostomocoris Schuh & Schwartz, 2004
 * Genus Agrametra Buchanan-White, 1878
 * Genus Arctostaphylocoris Schuh & Schwartz, 2004
 * Genus Atomophora Reuter, 1879
 * Genus Atomoscelis Reuter, 1875
 * Genus Atractotomoidea Yasunaga, 1999
 * Genus Atractotomus Fieber, 1858
 * Genus Badezorus Distant, 1910
 * Genus Beckocoris Knight, 1968
 * Genus Bergmiris Carvalho, 1984
 * Genus Boopidocoris Reuter, 1879
 * Genus Caiganga Carvalho & Becker, 1957
 * Genus Camptotylidea Wagner, 1957
 * Genus Campylomma Reuter, 1878
 * Genus Chinacapsus Wagner, 1961
 * Genus Chlamydatus Curtis, 1833
 * Genus Chlamyopsallus Schwartz, 2005
 * Genus Hambletoniola Carvalho, 1954
 * Genus Helenocoris Schmitz, 1976
 * Genus Hirtopsallus Schmitz, 1976
 * Genus Insulopus Schmitz, 1976
 * Genus Karocris V. Putshkov, 1975
 * Genus Kasumiphylus Schwartz & Stonedahl, 2004
 * Genus Knightomiroides Stonedahl & Schwartz, 1996
 * Genus Lamprosthenarus Poppius, 1914
 * Genus Larinocerus
 * Genus Lattinophylus Schuh, 2008
 * Genus Lindbergopsallus Wagner, 1962
 * Genus Lopsallus Schmitz, 1976
 * Genus Maurodactylus Reuter, 1878
 * Genus Megalopsallus Knight, 1927
 * Genus Monosynamma J. Scott, 1864
 * Genus Naresthus Schmitz, 1976
 * Genus Nasocoris Reuter, 1879
 * Genus Neisopsallus Schmitz, 1976
 * Genus Neophylus Carvalho & Costa, 1992
 * Genus Nevadocoris Knight, 1968
 * Genus Nigrimiris Carvalho & Schaffner, 1973
 * Genus Nigrocapillocoris Wagner, 1973
 * Genus Oligobiella Reuter, 1885
 * Genus Orthopidea Reuter, 1899
 * Genus Phaxia Kerzhner, 1984
 * Genus Phoenicocoris Reuter, 1875
 * Genus Pinomiris Stonedahl & Schwartz, 1996
 * Genus Pruneocoris Schuh & Schwartz, 2004
 * Genus Psallomimus Wagner, 1951
 * Genus Rhinacloa Reuter, 1876
 * Genus Salicarus Kerzhner, 1962
 * Genus Solenoxyphus Reuter, 1875
 * Genus Spanagonicus Berg, 1883
 * Genus Squamophylus Carvalho & Costa, 1992
 * Genus Sthenaropsis Poppius, 1912
 * Genus Taeniophorus Linnavuori, 1952
 * Genus Tannerocoris Knight, 1970
 * Genus Tapuruyunus Carvalho, 1946
 * Genus Thymopsallus Linnavuori, 1975
 * Genus Tijucaphylus Carvalho & Costa, 1992
 * Genus Tunisiella Carapezza, 1997
 * Genus Voruchia Reuter, 1879
 * Tribe Phylini Douglas & Scott, 1865
 * Subtribe Keltoniina Schuh & Menard, 2013
 * Genus Keltonia Knight, 1966
 * Genus Pseudatomoscelis Poppius, 1911
 * Genus Reuteroscopus Kirkaldy, 1905
 * Genus Waupsallus Linnavuori, 1975
 * Subtribe Oncotylina Douglas & Scott, 1865
 * Genus Acrotelus Reuter, 1885
 * Genus Alloeotarsus Reuter, 1885
 * Genus Americodema T. Henry, 1999
 * Genus Angelopsallus Schuh, 2006
 * Genus Antepia Seidenstiicker, 1962
 * Genus Arizonapsallus Schuh, 2006
 * Genus Asciodema Reuter, 1878
 * Genus Bisulcopsallus Schuh, 2006
 * Genus Brachyarthrum Fieber, 1858
 * Genus Cariniocoris T. Henry, 1989
 * Genus Ceratopsallus Schuh, 2006
 * Genus Cercocarpopsallus Schuh, 2006
 * Genus Chlorillus Kerzhner, 1962
 * Genus Compsidolon Reuter, 1899
 * Genus Crassomiris Weirauch, 2006
 * Genus Damioscea Reuter, 1883
 * Genus Dasycapsus Poppius, 1912
 * Genus Europiella Reuter, 1909
 * Genus Eurycolpus Reuter, 1875
 * Genus Galbinocoris Weirauch, 2006
 * Genus Glaucopterum Wagner, 1963
 * Genus Hamatophylus Weirauch, 2006
 * Genus Hoplomachus Fieber, 1858
 * Genus Ihermocoris Puton, 1875
 * Genus Insulaphylus Weirauch, 2006
 * Genus Josifovius Konstantinov, 2008
 * Genus Kmentophylus Duwal, Yasunaga, & Lee, 2010
 * Genus Knightophylinia Schaffner, 1978
 * Genus Knightopiella Schuh, 2004
 * Genus Knightopsallus Schuh, 2006
 * Genus Leucodellus Reuter, 1906
 * Genus Lineatopsallus T. Henry, 1991
 * Genus Litoxenus Reuter, 1885
 * Genus Maculamiris Weirauch, 2006
 * Genus Malacotes Reuter, 1878
 * Genus Marrubiocoris Wagner, 1970
 * Genus Megalocoleus Reuter, 1890
 * Genus Moiseevichia Schuh, 2006
 * Genus Nanopsallus Wagner, 1952
 * Genus Neopsallus Schuh & Schwartz, 2004
 * Genus Occidentodema T. Henry, 1999
 * Genus Oligotylus Van Duzee, 1916
 * Genus Omocoris Lindberg, 1930
 * Genus Oncotylidea Wagner, 1965
 * Genus Oncotylus Fieber, 1858
 * Genus Opisthotaenia Reuter, 1901
 * Genus Oreocapsus Linnavuori, 1975
 * Genus Parachlorillus Wagner, 1963
 * Genus Parapsallus Wagner, 1952
 * Genus Paredrocoris Reuter, 1878
 * Genus Phaeochiton Kerzhner, 1964
 * Genus Phallospinophylus Weirauch, 2006
 * Genus Phyllopidea Knight, 1919
 * Genus Phymatopsallus Knight
 * Genus Piceophylus Schwartz & Schuh, 1999
 * Genus Pinophylus Schwartz & Schuh, 1999
 * Genus Placochilus Fieber, 1858
 * Genus Plagiognathus Fieber, 1858
 * Genus Plesiodema Reuter, 1875
 * Genus Pleuroxonotus Reuter, 1903
 * Genus Pronototropis Reuter, 1879
 * Genus Psallodema V. Putshkov, 1970
 * Genus Psallomorpha Duwal Yasunaga, & Lee, 2010
 * Genus Psallovius T. Henry
 * Genus Pygovepres Weirauch, 2006
 * Genus Quercophylus Weirauch, 2006
 * Genus Quernocoris Weirauch, 2006
 * Genus Ranzovius Distant, 1893
 * Genus Rhinocapsus Uhler, 1890
 * Genus Roburocoris Weirauch, 2009
 * Genus Rubellomiris Weirauch, 2006
 * Genus Rubeospineus Weirauch, 2006
 * Genus Sacculifer Kerzhner, 1959
 * Genus Salicopsallus Schuh, 2006
 * Genus Schaffneropsallus Schuh, 2006
 * Genus Stenoparia Fieber, 1870
 * Genus Sthenaropsidea Henry & Schuh, 2002
 * Genus Stictopsallus Schuh, 2006
 * Genus Stirophylus Eckerlein & Wagner, 1965
 * Genus Stoebea Schuh, 1974
 * Genus Tapirula Carapezza, 1997
 * Genus Tinicephalus Fieber, 1858
 * Genus Tragiscocoris Fieber, 1861
 * Genus Vanduzeephylus Schuh & Schwartz, 2004
 * Genus Vesperocoris Weirauch, 2006
 * Genus Viscacoris Weirauch, 2009
 * Genus Zophocnemis Kerzhner, 1962
 * Subtribe Phylina Douglas & Scott, 1865
 * Genus Adelphophylus Wagner, 1959
 * Genus Agraptocoris Reuter, 1903
 * Genus Alnopsallus Duwal, Yasunaga, & Lee, 2010
 * Genus Alvarengamiris Carvalho, 1991
 * Genus Amazonophilus Carvalho & Costa, 1993
 * Genus Anapsallus Odhiambo, 1960
 * Genus Anomalocornis Carvalho & Wygodzinsky, 1945
 * Genus Arlemiris Carvalho, 1984
 * Genus Bicurvicoris Carvalho & Schaffner, 1973
 * Genus Botocudomiris Carvalho, 1979
 * Genus Brachycranella Reuter, 1905
 * Genus Conostethus Fieber, 1858
 * Genus Crassicornus Carvalho, 1945
 * Genus Darectagela Linnavuori, 1975
 * Genus Darfuromma Linnavuori, 1975
 * Genus Dignaia Linnavuori, 1975
 * Genus Dominiquella Linnavuori, 1983
 * Genus Ectagela Schmidt, K., 1939
 * Genus Ellacapsus Yasunaga, 2013
 * Genus Eremophylus Yasunaga, 2001
 * Genus Farsiana Linnavuori, 1998
 * Genus Gediocoris Wagner, 1964
 * Genus Ghazalocoris Linnavuori, 1975
 * Genus Gressittocapsus Schuh, 1984
 * Genus Icodema Reuter, 1875
 * Genus Indatractus Linnavuori, 1975
 * Genus Izyaius Schwartz, 2006
 * Genus Juniperia Linnavuori, 1965
 * Genus Knightensis Schaffner, 1978
 * Genus Lalyocoris Linnavuori, 1993
 * Genus Lasiolabopella Schuh, 1974
 * Genus Lepidargyrus Muminov, 1962
 * Genus Lepidocapsus Poppius, 1914
 * Genus Leptoxanthus Reuter, 1905
 * Genus Liviopsallus Carapezza, 1982
 * Genus Mendozaphylus Carvalho & Carpintero, 1991
 * Genus Millerimiris Carvalho, 1951
 * Genus Mixtecamiris Carvalho & Schaffner, 1973
 * Genus Natalophylus Schuh, 1974
 * Genus Nubaia Linnavuori, 1975
 * Genus Orthonotus Stephens, 1829
 * Genus Parafulvius Carvalho, 1954
 * Genus Paravoruchia Wagner, 1959
 * Genus Phylus Hahn, 1831
 * Genus Plagiognathidea Poppius, 1914
 * Genus Platyscytisca Costa & Henry, 1999
 * Genus Platyscytus Reuter, 1907
 * Genus Porophoroptera Carvalho & Gross, 1982
 * Genus Psallus Fieber, 1858
 * Genus Roudairea Puton & Reuter, 1886
 * Genus Sasajiophylus Yasunaga, 2001
 * Genus Somalocoris Linnavuori, 1975
 * Genus Sthenarus Fieber, 1858
 * Genus Stibaromma Odhiambo, 1961
 * Genus Tibiopilus Carvalho & Costa, 1993
 * Genus Trevessa China, 1924
 * Genus Villaverdea Carvalho, 1990
 * Genus Widdringtoniola Schuh, 1974
 * Genus Zakanocoris V. Putshkov, 1970
 * Genus Zanchiophylus Duwal, Yasunaga, & Lee, 2010
 * Tribe Pilophorini Douglas & Scott, 1876
 * Genus Alepidiella Poppius, 1914
 * Genus Aloea Linnavuori, 1975
 * Genus Dilatops Weirauch, 2006
 * Genus Druthmarus Distant, 1909
 * Genus Ethatractus Linnavuori, 1975
 * Genus Hypseloecus Reuter, 1891
 * Genus Lasiolabops Poppius, 1914
 * Genus Neoambonea Schuh, 1974
 * Genus Parambonea Schuh, 1974
 * Genus Parasthenaridea Miller, 1937
 * Genus Pherolepis Kulik, 1968
 * Genus Pilophorus Hahn, 1826
 * Genus Pseudambonea Schuh, 1974
 * Genus Randallophorus Henry, 2013
 * Genus Spinolosus Zou, 1985
 * Genus Sthenaridea Reuter, 1885
 * Tribe Semiini Knight, 1923
 * Subtribe Exocarpocorina Schuh & Menard, 2013
 * Genus Ampimpacoris Weirauch & Schuh, 2011
 * Genus Ancoraphylus Weirauch, 2007
 * Genus Araucanophylus Carvalho, 1984
 * Genus Basileobius Eyles & Schuh, 2003
 * Genus Chiloephylus Carvalho, 1984
 * Genus Cyrtodiridius Eyles & Schuh, 2003
 * Genus Exocarpocoris Weirauch, 2007
 * Genus Gonzalezinus Carvalho, 1981
 * Genus Halormus Eyles & Schuh, 2003
 * Genus Harpagophylus Schuh & Weirauch, 2010
 * Genus Jiwarli Soto & Weirauch, 2007
 * Genus Leptidolon Reuter, 1904
 * Genus Mecenopa Eyles & Schuh, 2003
 * Genus Melaleucoides Schuh & Weirauch, 2010
 * Genus Pimeleocoris Eyles & Schuh, 2003
 * Genus Polyozus Eyles & Schuh, 2003
 * Genus Thryptomenomiris Schuh & Weirauch, 2010
 * Genus Wallabicoris Schuh & Pedraza, 2010
 * Genus Xiphoidellus Weirauch & Schuh, 2011
 * Genus Xiphoides Eyles & Schuh, 2003
 * Subtribe Semiina Knight, 1923
 * Genus Criocoris Fieber, 1858
 * Genus Hoplomachidea Reuter, 1909
 * Genus Karoocapsus Schuh, 1974
 * Genus Monocris V. Putshkov, 1974
 * Genus Monospatha Eyles & Schuh 2003
 * Genus Myrmicopsella Poppius, 1914
 * Genus Nicholia Knight, 1929
 * Genus Parapseudosthenarus Schuh, 1974
 * Genus Pseudosthenarus Poppius, 1914
 * Genus Rakula Odhiambo, 1967
 * Genus Semium Reuter, 1876
 * Genus Thoth Linnavuori, 1993
 * Genus Tytthus Fieber, 1864