Ruderal species



A ruderal species is a plant species that is first to colonize disturbed lands. The disturbance may be natural – for example, wildfires or avalanches – or the consequences of human activities, such as construction (of roads, of buildings, mining, etc.) or agriculture (abandoned fields, irrigation, etc.).

The term ruderal originates from the Latin word rudus, meaning "rubble".

Ruderal species typically dominate the disturbed area for a few years, gradually losing the competition to other native species. However, in extreme disturbance circumstances, such as when the natural topsoil is covered with a foreign substance, a single-species ruderal community may become permanently established. In addition, some ruderal invasive species may have such a competitive advantage over the native species that they, too, may permanently prevent a disturbed area from returning to its original state despite natural topsoil.

Features
Features contributing to a species' success as ruderal are:
 * Massive seed production
 * Seedlings whose nutritional requirements are modest
 * Fast-growing roots
 * Independence of mycorrhizae
 * Polyploidy

Quantification
Ecologists have proposed various scales for quantifying ruderality, which can be defined as the "ability to thrive where there is disturbance through partial or total destruction of plant biomass" (Grime, Hodgson & Hunt, 1988). The ruderality scale of Grime presents values that are readily available, and it takes into account disturbance factors as well as other indicators such as the annual or perennial character of the plants.