Sclerosomatidae

The Sclerosomatidae are a family of harvestmen with about 1,300 known species. One former subfamily has been recently removed to form a new family, Globipedidae.

Name
The name is combined from Ancient Greek skleros "hard" and soma "body".

Genera

 * Gagrellinae Thorell, 1889
 * Abaetetuba Tourinho-Davis, 2004 (5 species)
 * Adungrella Roewer, 1955 (3 species)
 * Aguilaia González-Sponga, 2003 (1 species; Venezuela)
 * Akalpia Roewer, 1915 (2 species; India and Japan)
 * Altobunus Roewer, 1910 (3 species; Celebes and Philippines)
 * Amazonesia Soares, 1970 (2 species)
 * Antigrella Roewer, 1954 (1 species)
 * Aurivilliola Roewer, 1910 (17 species; South and Southeast Asia)
 * Azucarella Roewer, 1959 (1 species)
 * Bakerinulus Roewer, 1955 (1 species)
 * Bastia Roewer, 1910 (3 species; South and Southeast Asia)
 * Bastioides Mello-Leitão, 1931 (1 species; South America)
 * Baturitia Roewer, 1931 (1 species; Sunda Islands)
 * Bellonia González-Sponga, 2003 (1 species; Venezuela)
 * Biceropsis Roewer, 1935 (1 species; Burma)
 * Bonthainia Roewer, 1913 (5 species; South and Southeast Asia)
 * Bullobunus Roewer, 1910 (7 species; Philippines)
 * Caluga Roewer, 1959 (1 species)
 * Cardenalia Kury, 2017 (1 species; Venezuela)
 * Carinobius Roewer, 1955 (1 species)
 * Carmenia Roewer, 1915 (1 species; Colombia)
 * Carmichaelus Roewer, 1929 (1 species; India)
 * Ceratobunellus Roewer, 1911 (3 species; India)
 * Ceratobunoides Roewer, 1923 (2 species; Sumatra)
 * Ceratobunus Thorell, 1889 (13 species)
 * Cervibunus Roewer, 1912 (3 Species; South and Southeast Asia)
 * Chasenella Roewer, 1955 (2 species; Borneo)
 * Chebabius Roewer, 1935 (1 species; Burma)
 * Coonoora Roewer, 1929 (1 species; India)
 * Dentobunus Roewer, 1910 (33 species; Southeast Asia)
 * Diangathia Roewer, 1955 (1 species)
 * Echinobunus Roewer, 1912 (1 species; Java)
 * Euceratobunus Roewer, 1923 (1 species; India)
 * Eugagrella Roewer, 1910 (25 species; South and Southeast Asia)
 * Euzaleptus Roewer, 1911 (4 species; South and Southeast Asia)
 * Fesa Roewer, 1953 (1 species)
 * Gagrella Stoliczka, 1869 (226 species; Asia)
 * Gagrellenna Roewer, 1929 (1 species; India)
 * Gagrellina Roewer, 1913 (1 species; Celebes)
 * Gagrellissa Roewer, 1931 (1 species; Sunda Islands)
 * Gagrellopsis Sato & Suzuki, 1939 (1 species; Japan)
 * Gagrellula Roewer, 1910 (67 species; Asia)
 * Garleppa Roewer, 1912 (4 species; South America)
 * Geaya Roewer, 1910 (82 species; Latin America)
 * Globulosoma Martens, 1987 (2 species; Nepal)
 * Guaranobunus Ringuelet, 1954 (1 species; Argentina)
 * Hamitergum Crawford, 1992 (1 species; Russia)
 * Harmanda Roewer, 1910 (15 species; Asia)
 * Harmandina Schenkel, 1954 (1 species)
 * Hehoa Roewer, 1929 (1 species; India)
 * Heterogagrella Roewer, 1954 (2 species; Southeast Asia)
 * Hexazaleptus Suzuki, 1966 (1 species)
 * Hexomma Thorell, 1876 (9 species; Southeast Asia)
 * Himaldroma Martens, 1987 (2 species; Nepal)
 * Himalzaleptus Martens, 1987 (1 species; Nepal)
 * Holcobunus Roewer, 1910 (26 or 27 species; Latin America)
 * Holmbergiana Mello-Leitão, 1931 (3 species; South America)
 * Hologagrella Roewer, 1910 (7 species; Southeast Asia)
 * Hypogrella Roewer, 1955 (1 species)
 * Hypsibunus Thorell, 1891 (11 species; India & Southeast Asia)
 * Jussara Mello-Leitão, 1935 (10 species; South America)
 * Koyamaia Suzuki, 1972 (1 species)
 * Krusa Goodnight & Goodnight, 1947 (10 species; Mexico)
 * Krusella Roewer, 1953 (1 species; Venezuela)
 * Liopagus Chamberlin, 1916 (1 species)
 * Marthana Thorell, 1891 (24 species; Southeast Asia)
 * Melanopa Thorell, 1889 (30 species; India, Southeast Asia)
 * Melanopella Roewer, 1931 (3 species; Sunda Islands)
 * Melanopula Roewer, 1929 (5 species; South and Southeast Asia)
 * Metadentobunus Roewer, 1915 (3 species; Taiwan)
 * Metahehoa Suzuki, 1985 (1 species)
 * Metasyleus Roewer, 1955 (3 species)
 * Metaverpulus Roewer, 1912 (7 species; South Asia)
 * Metazaleptus Roewer, 1912 (8 species; South and Southeast Asia)
 * Microzaleptus Roewer, 1955 (1 species)
 * Munequita Mello-Leitão, 1941 (1 species; Brazil)
 * Neogagrella Roewer, 1913 (3 species; South and Southeast Asia)
 * Nepalgrella Martens, 1987 (2 species; Nepal)
 * Nepalkanchia Martens, 1990 (2 species; Nepal)
 * Obigrella Roewer, 1955 (1 species)
 * Octozaleptus Suzuki, 1966 (1 species)
 * Onostemma Mello-Leitão, 1938 (2 species; Brazil)
 * Oobunus Kishida, 1930 (1 species)
 * Orissula Roewer, 1955 (1 species)
 * Padangrella Roewer, 1954 (1 species)
 * Palniella Roewer, 1929 (1 species; India)
 * Paradentobunus Roewer, 1915 (1 species; India)
 * Paragagrella Roewer, 1912 (5 species; South and Southeast Asia)
 * Paragagrellina Schenkel, 1963 (1 species)
 * Parageaya Mello-Leitão, 1933 (6 species; Latin America)
 * Paratamboicus Mello-Leitão, 1940 (30 species; South America)
 * Paraumbogrella Suzuki, 1963 (1 species)
 * Paruleptes Soares, 1970 (1 species)
 * Pectenobunus Roewer, 1910 (4 species; South America)
 * Pergagrella Roewer, 1954 (1 species)
 * Pokhara Suzuki, 1970 (7 species; Nepal)
 * Prionostemma Pocock, 1903 (114 species; Mexico, Central America, and South America)
 * Prodentobunus Roewer, 1923 (4 species; Asia)
 * Psammogeaya Mello-Leitão, 1946 (1 species; Uruguay)
 * Psathyropus L. Koch, 1878 (40 species; Asia)
 * Pseudarthromerus Karsch, 1892 (1 species)
 * Pseudogagrella Redikortsev, 1936 (13 species; East Asia)
 * Pseudomelanopa Suzuki, 1974 (2 species; East Asia)
 * Pseudosystenocentrus Suzuki, 1985 (1 species)
 * Romerella Goodnight & Goodnight, 1943 (5 species; Mexico and South America)
 * Sarasinia Roewer, 1913 (1 species; Celebes)
 * Sataria Roewer, 1915 (3 species; India)
 * Scotomenia Thorell, 1889 (1 species; Burma)
 * Sericicorpus Martens, 1987 (1 species; Nepal)
 * Sinadroma Roewer, 1955 (1 species)
 * Syleus Thorell, 1876 (2 species; India)
 * Syngagrella Roewer, 1913 (1 species; Celebes)
 * Systenocentrus Simon, 1886 (6 species; Southeast and East Asia)
 * Tamboicus Roewer, 1912 (3 species; South America)
 * Taperina Roewer, 1953 (2 species)
 * Tetraceratobunus Roewer, 1915 (3 species; South Asia)
 * Thunbergia Martens, 2020 (4 species; South America)
 * Toragrella Roewer, 1955 (2 species)
 * Trachyrhinus Weed, 1892 (7 species; North America: Mexico and United States)
 * Umbogrella Roewer, 1955 (1 species)
 * Umbopilio Roewer, 1956 (2 species)
 * Vaho González-Sponga, 2003 (1 species)
 * Valle González-Sponga, 2003 (3 species)
 * Varinodulia Canals, 1935 (1 species)
 * Verpulus Simon, 1901 (13 species; South and Southeast Asia)
 * Verrucobunus Roewer, 1931 (3 species; East and Southeast Asia)
 * Xerogrella Martens, 1987 (1 species; Nepal)
 * Zaleptiolus Roewer, 1955 (4 species; Asia)
 * Zaleptulus Roewer, 1955 (3 species)
 * Zaleptus Thorell, 1876 (72 species; Asia and Australia)


 * Gyantinae Silhavý, 1946
 * Gyas Simon, 1879 (1 species; Alps)
 * Gyoides Martens, 1982 (6 species; Nepal)
 * Rongsharia Roewer, 1957 (3 species; Nepal)


 * Leiobuninae Banks, 1893
 * †Amauropilio Mello-Leitão, 1937 (fossil) (2 species; fossils found in Colorado)
 * Cosmobunus Simon, 1879 (5 species; Spain and Mexico)
 * Dilophiocara Redikorzev, 1931 (2 species)
 * Eumesosoma Cokendolpher, 1980 (6 species; United States)
 * Eusclera Roewer, 1910 (2 species; China and India)
 * Hadrobunus Banks, 1900 (2 species; United States)
 * Leiobunum C.L.Koch, 1839 (126 species - 2 extinct fossils; worldwide)
 * Leuronychus Banks, 1900 (2 species; North America)
 * Microliobunum Roewer, 1912 (2 species; Middle East)
 * Micronelima Schenkel, 1938 (1 species; Spain)
 * Nelima Roewer, 1910 (44 species; worldwide)
 * Paranelima Caporiacco, 1938 (6 species; Mexico and Central America)
 * Schenkeliobunum Starega, 1964 (1 species)
 * Togwoteeus Roewer, 1952 (1 species; United States)


 * Sclerosomatinae Simon, 1879
 * Astrobunus Thorell, 1876 (12 species; Europe)
 * Granulosoma Martens, 1973 (1 species)
 * Homalenotus C.L.Koch, 1839 (13 species; Southern Europe and North Africa)
 * Mastobunus Simon, 1879 (2 species; Southern Europe and North Africa)
 * Metasclerosoma Roewer, 1912 (5 species; Italy)
 * Pseudastrobunus Martens, 1973 (1 species)
 * Pseudohomalenotus Caporiacco, 1935 (1 species; the Karakoram)
 * Pygobunus Roewer, 1957 (2 species; Japan)
 * Umbopilio Roewer, 1956 (1 species)

Removed

 * Caiza Roewer, 1925 . Subjective synonym of Pectenobunus Roewer, 1910) in Tourinho, 2004a


 * Pseudoarthromerus Karsch, 1891 (1 species; Japan, nomen nudum, therefore non-valid)