Scutiger (frog)

Scutiger is a genus of toads in the family Megophryidae. Common name lazy toads has been coined for them. They occur in China, Burma, Nepal, and northern India in high-altitude habitats. Most are endemic to China.

Evolution
A 2017 molecular phylogenetic study found that Scutiger originated in Paleo-Tibet during the Oligocene.

Species
The following species are recognised in the genus Scutiger:
 * Scutiger adungensis Dubois, 1979 – Adung lazy toad
 * Scutiger bhutanensis Delorme and Dubois, 2001
 * Scutiger boulengeri (Bedriaga, 1898) – Boulenger's lazy toad, Xizang alpine toad, Himalayan stream frog
 * Scutiger brevipes (Liu, 1950)
 * Scutiger chintingensis Liu and Hu, 1960 – Chinting lazy toad, Chinting alpine toad
 * Scutiger ghunsa Khatiwada, Shu, Subedi, Wang, Ohler, Cannatella, Xie, and Jiang, 2019 – Ghunsa high altitude toad
 * Scutiger glandulatus (Liu, 1950) – Hopachai lazy toad, chest gland cat-eyed toad
 * Scutiger gongshanensis Yang and Su, 1979 – Gongshan lazy toad, Gongshan cat-eyed toad
 * Scutiger jiulongensis Fei, Ye, and Jiang, 1995 – Jiulong cat-eyed toad
 * Scutiger liupanensis Huang, 1985 – Liupan lazy toad, Liupan alpine toad
 * Scutiger maculatus (Liu, 1950) – spotted lazy toad, piebald alpine toad
 * Scutiger mammatus (Günther, 1896) – Tungsolo lazy toad
 * Scutiger muliensis Fei and Ye, 1986 – Muli cat-eyed toad
 * Scutiger nepalensis Dubois, 1974 – Nepal lazy toad, mountain pelobatid toad, Asiatic spadefoot toad
 * Scutiger ningshanensis Fang, 1985 – Ningshan lazy toad, Ningshan alpine toad
 * Scutiger nyingchiensis Fei, 1977 – Nyingchi lazy toad, Ladakh pelobatid toad, Nyingshi alpine toad
 * Scutiger occidentalis Dubois, 1978 – Ladakh high altitude toad
 * Scutiger pingwuensis Liu and Tian, 1978 – Pingwu lazy toad, Pingwu alpine toad
 * Scutiger sikimmensis (Blyth, 1855) – Sikkim lazy toad, Sikkim high altitude toad, Blyth's short-limbed frog
 * Scutiger spinosus Jiang, Wang, Li, and Che, 2016 – spiny lazy toad
 * Scutiger tengchongensis Yang & Huang, 2019
 * Scutiger tuberculatus Liu and Fei, 1979 – bumpy lazy toad, round-tubercled cat-eyed toad
 * Scutiger wanglangensis Ye and Fei, 2007 – Wanglang alpine toad
 * Scutiger wuguanfui Jiang, Rao, Yuan, Wang, Li, Hou, Che, and Che, 2012 – Wu's lazy toad, Medog lazy toad

Endemic ranges
Many Scutiger species are endemic to highly restricted geographical areas in the Eastern Himalayas. The ranges often overlap with those of Oreolalax species.


 * Ningxia, China
 * Scutiger liupanensis: Liupanshan National Nature Reserve, Jingyuan County, Ningxia Hui Autonomous Region, China
 * Shaanxi, China
 * Scutiger ningshanensis: Ningshan County, Shaanxi
 * Sichuan, China
 * Scutiger brevipes: Dawu County, Sichuan
 * Scutiger chintingensis: Mount Emei area, Sichuan
 * Scutiger jiulongensis: Jiulong County, Sichuan
 * Scutiger maculatus: Garze, Sichuan and Jiangda, Tibet
 * Scutiger muliensis: Muli County, Sichuan
 * Scutiger pingwuensis: Pingwu County, Sichuan
 * Scutiger wanglangensis: Sichuan
 * Yunnan, China
 * Scutiger gongshanensis: Gongshan County, Yunnan
 * Tibet
 * Scutiger maculatus: Garze, Sichuan and Jiangda, Tibet
 * Scutiger nyingchiensis: Nyingchi Prefecture, Tibet
 * Scutiger spinosus: Medog County, Tibet and Tawang District, Arunachal Pradesh
 * Scutiger wuguanfui: Medog County, Tibet
 * Bhutan
 * Scutiger bhutanensis: Bhutan
 * Sikkim, India
 * Scutiger sikimmensis: Sikkim, India