Tlayúa Formation

The Tlayúa Formation is an Early Cretaceous (late Albian) geological formation near Tepexi de Rodríguez, Puebla.

Paleobiota
The formation contains a diverse array of vertebrate and invertebrate fossils. About 70% of the macrofossils are osteichthyan fish. Other vertebrates include chelonians, pterosaurs, lepidosaurs, and crocodiles. Cyanobacteria, foraminifera, algae, gymnosperms, sponges, cnidarians, annelids, gastropods, ammonites, bivalves, arachnids, insects, isopods, anomurans, brachyurans, crinoids, echinoids, holothuroids, stelleroids, and ophiuroids, have also been recovered from the Tlayúa Formation.

Fish
Mainly after González-Rodríguez (2016) and Applegate et al. (2006)


 * Araripichthys webri
 * Armigatus carrenoae
 * Armigatus felixi
 * Axelrodichthys cf. araripensis
 * Cipactlichthys scutatus
 * Epaelops martinezi
 * Macrosemiocotzus americanus
 * Michin csernai
 * Notagogus novomundi
 * Nusaviichthys nerivelai
 * Pachyamia mexicana
 * Paraclupea seilacheri
 * Quetzalichthys perrilliatae
 * Ranulfoichthys dorsonudum
 * Teoichthys brevipina
 * Teoichthys kallistos
 * Tepexichthys aranguthyrom
 * Tlalocbatos applegatei
 * Tlayuamichin itztli
 * Unamichthys espinosai
 * Ellimmichthys sp.
 * cf. Amblysemius
 * cf. Bananogmius
 * Belonostomus sp.
 * cf. Brannerion
 * Lepidotes sp.
 * cf. Lycoptera
 * cf. Megalops
 * Neoproscinetes sp.
 * Notelops sp.
 * cf. Paraelops
 * cf. Rhacolepis
 * Vinctifer sp.
 * cf. Yabrudichthys
 * Gonorhynchiform indet.

Other vertebrates

 * Testudines indet. (at least two taxa)
 * Pterosauria indet. (tentatively assigned as Nyctosaurus and Pteranodon)
 * Crododylia indet.

Arthropods

 * Archaeoniscus aranguthyorum
 * Atocatle ranulfoi
 * Ixtahua benjamini
 * Protaegla miniscula
 * Tepexicarcinus tlayuaensis
 * Sphaeromatidae sp.
 * Ostracoda?

Mollusks

 * Inoceramus sp.
 * Pectinidae indet.
 * Osteridae indet.
 * Gastropoda indet.
 * Neohibolites minimus obtusus
 * Neohibolites minimus pinguis
 * Neohibolites minimus claviformis
 * Neohibolites praeultimus
 * Mesohibolites semicanaliculatus
 * Martoniceras sp.
 * Hysteroceras sp.
 * Anisoceras sp.

Echinoderms

 * Ophiactis applegatei
 * Paleopentacta alencasterae
 * Parapsolus tlayuensis
 * Astropecten sp.
 * Echinaster sp.
 * Ophidiaster sp.
 * Plutonaster sp.
 * Tamaria sp.
 * Crinoidea indet.

Other invertebrates

 * Porifera
 * Anthozoa
 * Annelida

Plants

 * cf. Araucaria sp.
 * cf. Frenolopsis
 * Brachyphyllum sp.
 * Podozamites sp.
 * Zamites sp.
 * Phaeophyceae