Tylototriton

Tylototriton is a genus of newts known as crocodile newts or knobby newts. About 38 known species are in this genus. Many species have been described just recently. They range from northeastern India and Nepal through Burma to northern Thailand, Laos, Vietnam, and southern China.

Species
38 species recognized as of July 2023:


 * Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015
 * Tylototriton anhuiensis Qian, Sun, Li, Guo, Pan, Kang, Wang, Jiang, Wu, and Zhang, 2017
 * Tylototriton asperrimus Unterstein, 1930
 * Tylototriton broadoridgus Shen, Jiang, and Mo, 2012
 * Tylototriton dabienicus Chen, Wang, and Tao, 2010
 * Tylototriton daloushanensis Zhou, Xiao, and Luo, 2022
 * Tylototriton hainanensis Fei, Ye, and Yang, 1984
 * Tylototriton himalayanus Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015
 * Tylototriton kachinorum Zaw, Lay, Pawangkhanant, Gorin, and Poyarkov, 2019
 * Tylototriton kweichowensis Fang and Chang, 1932
 * Tylototriton liuyangensis Yang, Jiang, Shen, and Fei, 2014
 * Tylototriton lizhengchangi Hou, Zhang, Jiang, Li and Lu, 2012
 * Tylototriton maolanensis Li, Wei, Cheng, Zhang, and Wang, 2020
 * Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018
 * Tylototriton ngoclinhensis Phung, Pham, Nguyen, Ninh, Nguyen, Bernardes, Le, Ziegler & Nguyen, 2023
 * Tylototriton notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010
 * Tylototriton panhai Nishikawa, Khonsue, Pomchote, and Matsui, 2013
 * Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019
 * Tylototriton pasmansi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
 * Tylototriton phukhaensis Pomchote, Khonsue, Thammachoti, Hernandez, Peerachidacho, Suwannapoom, Onishi, and Nishikawa, 2020
 * Tylototriton podichthys Phimmachak, Aowphol, and Stuart, 2015
 * Tylototriton pseudoverrucosus Hou, Gu, Zhang, Zeng, and Lu, 2012
 * Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012
 * Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995
 * Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014
 * Tylototriton sini Lyu, Wang, Zeng, Zhou, Qi, Wan, and Wang, 2021
 * Tylototriton sparreboomi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
 * Tylototriton taliangensis Liu, 1950
 * Tylototriton thaiorum Poyarkov, Nguyen, and Arkhipov, 2021
 * Tylototriton tongziensis Li, Liu, Shi, Wei, and Wang, 2022
 * Tylototriton umphangensis Pomchote, Peerachidacho, Hernandez, Sapewisut, Khonsue, Thammachoti, and Nishikawa, 2021
 * Tylototriton uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013
 * Tylototriton verrucosus Anderson, 1871
 * Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005
 * Tylototriton wenxianensis Fei, Ye, and Yang, 1984
 * Tylototriton yangi Hou, Zhang, Zhou, Li, and Lu, 2012
 * Tylototriton zaimeng Decemson, Lalremsanga, Elangbam, Vabeiryureilai, Shinde, Purkayastha, Arkhipov, Bragin & Poyarkov, 2023
 * Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013

Phylogeny
The following phylogeny of Tylototriton is from Nishikawa, Matsui & Rao (2014).