Urocyclidae

Urocyclidae is a family of air-breathing land snails, semi-slugs and land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea, (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Some species within this family make and use love darts before mating.

Distribution
The Urocyclidae are distributed in the Afrotropical realm, including Madagascar, the Comores and the islands of the Gulf of Guinea.

Taxonomy
The family Urocyclidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005), that follows classification of Urocyclinae by Van Goethem (1977):
 * subfamily Rhysotininae Schileyko, 2002
 * subfamily Sheldoniinae Connolly, 1925 (1912) - synonyms: Peltatinae Godwin-Austen, 1912; Trochonanininae Connolly, 1912; Trochozonitinae Iredale, 1914; Ledoulxiinae Pilsbry, 1919; Gymnarionidae Von Mol, 1970; Rhysotinidae Schileyko, 2002; Zonitarionini Schileyko, 2002; Acantharionini Schileyko, 2002
 * subfamily Urocyclinae Simroth, 1889
 * tribe Urocyclini Simroth, 1889 - synonyms: Atoxonini Schileyko, 2002; Buettneriini Schileyko, 2002
 * tribe Dendrolimacini Van Goethem, 1977
 * tribe Leptichnini Van Goethem, 1977
 * tribe Upembellini Van Goethem, 1977

Genera
Genera within the family Urocyclidae include:
 * unsorted:
 * † Koruella Pickford, 2009


 * Subfamily Rhysotininae Schileyko, 2002
 * Rhysotina Ancey, 1887
 * Thomeonanina Germain, 1909 : synonym of Rhysotina Ancey, 1887 (objective synonym)
 * Subfamily Sheldoniinae
 * Acantharion Binder & Tillier, 1985
 * Africarion Godwin-Austen, 1883
 * Amatarion Van Mol, 1970
 * Angustivestis Pilsbry, 1919
 * Aspidelus Morelet, 1884
 * Aspidotomium Degner, 1932
 * Belonarion Pilsbry, 1919
 * Bloyetia Bourguignat, 1890
 * Camerunarion Van Mol, 1970
 * Carinazingis van Bruggen & de Winter, 1990
 * Centrafricarion Van Mol, 1970
 * Chlamydarion Van Mol, 1968
 * Colparion Laidlaw, 1938
 * Degneria Verdcourt, 1956
 * Entagaricus Pilsbry, 1919
 * Estria Poirier, 1887
 * Falloonella Preston, 1914
 * Fuellebornia Verdcourt, 1998
 * Granularion Germain, 1912
 * Gymnarion Pilsbry, 1919
 * Hamya Bourguignat, 1885 †
 * Kerkophorus Godwin-Austen, 1912
 * Lacrimarion Connolly, 1939
 * Malagarion Tillier, 1979
 * Mesafricarion Pilsbry, 1919
 * Microkerkus Godwin-Austen, 1912
 * Montanobloyetia Verdcourt, 1961
 * Morrumbalia Verdcourt, 1998
 * Paragranularion Van Mol, 1970
 * Percivalia Preston, 1914
 * Plicatonanina Verdcourt, 1961
 * Principicochlea D. Holyoak & G. Holyoak, 2020
 * Principitrochoidea D. Holyoak & G. Holyoak, 2020
 * Ptilototheca Herbert, 2016
 * Rhopalogonium Degner, 1932
 * Selatodryas Herbert, 2017
 * Senegalarion Van Mol, 1970
 * Sheldonia Ancey, 1887 - the type genus of the subfamily Sheldoniinae
 * Sjostedtina Verdcourt, 1961
 * Sylvarion Van Mol, 1970
 * Thapsia Albers, 1860
 * Thielarion Van Mol, 1970
 * Thomithapsia D. Holyoak & G. Holyoak, 2020
 * Thomitrochoidea D. Holyoak & G. Holyoak, 2020
 * Tresia Van Goethem, 1975
 * Trochonanina Mousson, 1869
 * Trochozonites Pfeffer, 1883
 * Tropidocochlion Verdcourt, 1998
 * Verdcourtia Van Mol, 1970
 * Verrucarion Van Mol, 1970
 * Zingis E. von Martens, 1878
 * Zonitarion Pfeffer, 1883
 * Ledoulxia Bourguignat, 1885: synonym of Trochonanina Mousson, 1869
 * Peltatus Godwin-Austen, 1908: synonym of Sheldonia Ancey, 1887

Tribe Dendrolimacini
 * Subfamily Urocyclinae
 * Dendrolimax Heynemann, 1868 - the type genus of the tribe Dendrolimacini

Tribe Leptichnini
 * Leptichnus Simroth, 1896 - the type genus of the tribe Leptichnini

Tribe Upembellini
 * Leptichnoides Van Goethem, 1975
 * Upembella Van Goethem, 1969 - the type genus of the tribe Upembellini

tribe Urocyclini
 * Anisotoxon Van Goethem, 1975
 * Atoxon Simroth, 1888
 * Atoxonoides Van Goethem, 1973
 * Atrichotoxon Simroth, 1910
 * Buettneria Simroth, 1910
 * Bukobia Simroth, 1896
 * Elisolimax Cockerell, 1893
 * Emphysetes Verdcourt, 2003
 * Microcyclus Simroth, 1896
 * Nupnus Van Goethem, 1975
 * Pembatoxon Van Goethem, 1975
 * Phaneroporus Simroth, 1888
 * Polytoxon Simroth, 1897
 * Pseudatoxon Van Goethem, 1975
 * Tanzalimax Rowson, Paustian & Van Goethem, 2017
 * Trichotoxon Simroth, 1888
 * Udzungwalimax Rowson, Paustian & Van Goethem, 2017
 * Urocyclus Gray, 1864 - the type genus of the family Urocyclidae

Cladogram
The following cladogram shows the phylogenic relationship of this family to other families in the limacoid clade: