User:DavidAnstiss/Ignurbia

Ignurbia is a monotypic genus of flowering plants in the daisy family. There is only one known species, Ignurbia constanzae, which is native to the island of Hispaniola in the West Indies (Haiti and Dominican Republic).

Description
It is a perennial herb, 3 –

''Erect, little-branched, herbaceous, suffrutescent herb, 0.5-2 m high, sparsely pubescent on stems, leaves and peduncles; trichomes unbranched, mostly multicellular, articulated, tapering from a wider base, white or brownish, some gland-tipped, some slender, unicellular. Leaves alternate, petiolate, ovate to oblong-obovate in outline, 5-14 × 4-9 cm, irregularly pinnately 3-5-lobed or pinnatifid, apically acute, basally subtruncate or somewhat cuneately tapering into the petiole, herbaceous, pinnately veined with reticulately anastomosing veinlets, laxly hirsute, leaf margins laxly dentate with mucronulate tips; petiole 2-6 cm long. Synflorescence terminal, pedunculate, densely corymbose; peduncles 6-25 cm long. Capitula numerous, homogamous, discoid, 15- 21-flowered, calyculate. Involucre cupshaped-cylindrical; involucral bracts 8-13, ± uniseriate, linear-lanceolate, 7-10 × 0.7-1.5 mm, herbaceous with scarious, whitish margins, acute to acuminate, with 3-5 resiniferous veins, glabrous except for a few scattered short setae and distinctly puberulous tips; calyculus bracts 4-6, linear, 2 mm long. Receptacle flat or slightly convex, glabrous, minutely alveolate. Corolla orange or dull to dirty yellow, 7-10 mm long, tubular and gradually widening above; lobes lanceolate or narrowly ovate, 1.5-2 mm long, glabrous, faintly midlined and with distinct lateral veins continuing down the corolla, margins involute, apex acute. Stamens and styles much exserted. Anthers 2.5-3 mm long, basally obtuse and shortly auriculate; apical appendage oblong-lanceolate, obtuse; endothecial tissue radial with short, almost isodiametric cells with thickenings on all walls; filament collar long, + uniformly wide but with enlarged marginal cells. Pollen grains tricolporate, very minutely spinulose. Style branches linear, c. 2 mm long, with continuous stigmatic areas inside, apically obtuse to rounded without distinct sweeping-hairs and just with a few short pili or papillae, dorsal side shortly glandular-papillate in the distal half. Cypselas narrowly oblong, subterete but some subcompressed or slightly triquetrous, 2-3 × 0.7-0.8 mm, glabrous, dark brown, with 10 prominent honey-coloured ribs; carpopodium distinct, of 6-7 cell layers; ovary wall crystals small, non-prismatic, like sand. Pappus bristles pluriseriate, slender, only minutely barbellate, white, 6-8 mm long, basally connate, persistent.''.

The new genus is characterized by its herbaceous, suffruticose habit, dissected leaves with a herbaceous texture, orange or dirty yellow discoid capitula and styles with continuous stigmatic areas lacking distinct sweeping-hairs.

Taxonomy
It was originally published as Senecio constanzae by Ignatz Urban in Symb. Antill. vol.7 on page 430 in 1912. Ignurbia constanzae was found to be distinct from other Senecio species due to various features such as disc floret styles, stamens, endothecial tissue of the anthers and pollen grain shape. Then the genus, Ignurbia and species, Ignurbia constanzae were both first published and described by Swedish botanist Bertil Nordenstam in Willdenowia vol.36 on page 464 in 2006.

The genus name of Ignurbia is in honour of Ignatz Urban (1848–1931), who was a German botanist. He is known for his contributions to the flora of the Caribbean and Brazil, and for his work as curator of the Berlin Botanical Garden.

The specific epithet of constanzae refers to the town of Constanza near the Dominican city of Santo Domingo.

It is possibly related to genera in tribe Senecioneae; such as Odontocline (found in Jamaica), Jessea (in Costa Rica and Panama) and Talamancalia (in Costa Rica, Panama, Ecuador and Peru).

Habitat
Ignurbia constanzae grows in montane, humid forest or broad-leaved scrub with, e.g., Brunellia comocladifolia, Garrya fadyenii and sometimes Pinus occidentalis, and along streams and brooks, at altitudes from 1250 to 2400 m. 3 –