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Tufted Duck
The Tufted Duck dives for fifteen seconds to find food.

Early life
Casselton was born on 18 July 1938 to William Charles Henry Smith and Cecile Smith (née Bowman) in Rochford, Essex. Her parent's smallholding and her father's interest in natural history and genetics encouraged her and her sister Pauline in the direction of biology. She was educated at Southend High School for Girls, a grammar school in Southend-on-Sea. She studied at University College London, from which she gained a Bachelor of Science (BSc) degree in botany and a Doctor of Philosophy (PhD) degree in 1964.

Academic career
Casselton began her career in lecturing and research as an assistant lecturer at Royal Holloway College in London. She was Professor of Genetics at Queen Mary University of London from 1989–91 and was later awarded an AFRC/BBSRC Postdoctoral Fellowship, followed by a BBSRC Senior Research Fellowship in 1995.

Casselton was a Fellow of St Cross College Oxford from 1993–2003, and was appointed Professor of Fungal Genetics at Oxford in 1997. Her specialism was sexual development in fungi and she contributed to over 100 publications on this topic. She was a Fellow of St Cross College, Oxford from 1993 to 2003, and an Honorary Fellow of St Hilda's College, Oxford from 2000. She was a member of the Royal Society's Council from 2002–2003, and rejoined the Council in 2006 as Vice President and Foreign Secretary, replacing Professor Dame Julia Higgins.

As Foreign Secretary of the Royal Society, Casselton gave the Royal Society Rutherford Lecture in South Africa and the Blackett Lecture in India, travelling to 27 different countries during three and a half years in office.

Personal life
She married Peter John Casselton in 1961, divorcing him in 1978. She married William Joseph Dennis Tollett in 1981.

Awards and Honours
She was elected a Fellow of the Royal Society in 1999. She became a Member of the Academia Europaea in 2008, and was awarded an Honorary Doctor of Science by Queen Mary College, University of London in 2009 and University College London in September 2010. She was appointed Commander of the Order of the British Empire (CBE) in the 2012 Birthday Honours for services to fungal genetics and international science. Her nomination for the Royal Society reads:

The British Mycological Society awarded her an Honorary Membership in 2002.

Selected Publications

 * Stajich, JE., and 49 others, "Insights into evolution of multicellular fungi from the assembled chromosomes of the mushroom Coprinopsis cinerea (Coprinus cinereus). " Proceedings of the National Academy of Sciences (USA), Vol 107 (2010) pp. 11889–11894
 * Riquelme, M., Challen, MP., Casselton, LA., Brown, AJ. "The origin of multiple B mating specificities in Coprinus cinereus. " Genetics, Vol 170 (2005) pp. 1105–1119
 * Casselton, LA. "Mate recognition in fungi." Heredity, Vol 88 (2002) pp. 142–147
 * Brown, AJ, Casselton, LA. "Mating in mushrooms: increasing the chances but prolonging the affair." Trends in Genetics, Vol 17 (2001) pp. 393–400
 * Olesnicky, NS., Brown, AJ., Dowell, SJ., Casselton, LA. "A constitutively active G-protein-coupled receptor causes mating self-compatibility in the mushroom Coprinus" EMBO Journal, Vol 18 (1999) pp. 2756–2763
 * Casselton, LA., Olesnicky, NS. "Molecular genetics of mating recognition in basidiomycete fungi." Microbiology and Molecular Biology Reviews, Vol 62 (1998) pp. 55-
 * Banham, AH., Asanteowusu, RN., .... Casselton, LA. "An N-terminal dimerization domain permits homeodomain proteins to choose compatible partners and initiate sexual development in the mushroom Coprinus cinereus." Plant Cell, Vol 7 (1995) pp. 773–783
 * Kues, U., Casselton, LA. "Fungal mating type genes - regulators of sexual development." Mycological Research, Vol 96 (1992) pp. 993–1006
 * Kues, U., Richardson, WVJ., ... Casselton, LA. "The combination of dissimilar alleles of the A-alpha and A-beta gene complexes, whose proteins contain homeo domain motifs, determines sexual development in the mushroom Coprinus cinereus." Genes and Development, Vol 6 (1992) pp. 568–577
 * Binninger, DM., Skrzynia, C., Pukkila, PJ., Casselton, LA. "DNA-mediated transformation of the basidiomycete Coprinus cinereus." EMBO Journal, Vol 6 (1987) pp. 835–840
 * Sealy-Lewis, HM., Casselton, LA. "Restoration of enzyme activity by recessive missense suppressors in fungus Coprinus." Molecular and General Genetics, Vol. 164 (1978) pp. 211–215
 * Casselton, LA., Lewis, D. "Dilution of gene products in cytoplasm of heterokaryons in Coprinus lagopus". Genetical Research, Vol. 9, pp. 63-
 * Casselton, LA. "Production and behaviour of diploids of Coprinus lagopus." Genetical Research, Vol. 6 (1965) pp. 190-