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Moseley was born in the Cotswolds village of Sheepscombe, Gloucestershire, England. William has two siblings (a brother and a sister), which are both younger than him. Daisy Elizabeth Moseley was born in 1989 and Benjamin Hugh Moseley was born in 1992. He is the son of Juliette E. (née Fleming) and Peter A. Moseley, a cinematographer. Moseley attended Sheepscombe Primary School from September 1991 to July 1998, and then continued his education at Wycliffe College, also taking time off at Downfield Sixth Form, Stroud. Moseley began dating actress Kelsey Chow in 2012, his co-star in the film Run.

In 2013, Moseley starred as Daniel Lombardi in the action thriller film Run alongside Kelsey Chow. In 2014, Moseley played Anderl Gruber in the adventure drama The Silent Mountain, which follows a young Austrian soldier in World War I who fights his way through the Alps to rescue his first love and escape the impending explosion that will rock the mountain. While shooting this film, The Silent Mountain, William was badly electrocuted due to a lightning storm that occured on set. As a result, he has a scar on his bicep.