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Origins and name of The Bronx
The Bronx was called Rananchqua by the native Siwanoy band of Lenape (the Delawares to Europeans), while other Native Americans knew the Bronx as Keskeskeck. It was divided by the Aquahung River.

Jonas Bronck (ca. 1600–1649), a Swedish sea-captain working for the Dutch, entering New Netherland in 1639, became the first recorded European settler in the area. He leased land from the Dutch West India Company on the neck of the mainland immediately north of the Dutch settlement in Harlem (on Manhattan island), and bought additional tracts from the local tribes. He eventually accumulated 500 acres (about 2 square km, or 3/4 of a square mile) between the Harlem River and the Aquahung, which became known as Bronck's River, or The Bronx. Dutch and English settlers referred to the area as Bronck's Land. The American poet William Bronk is a descendant of Pieter Bronck, either Jonas Bronck's son or younger brother.

The Bronx is referred to, both legally, and colloquially, with a definite article, as The Bronx. (The County of Bronx, unlike the coextensive Borough of the Bronx, does not place the immediately before Bronx in formal references, nor does the United States Postal Service in its database of Bronx addresses.) The name for this region, apparently after the Bronx River, first appeared in the Annexed District of the Bronx created in 1874 out of part of Westchester County and was continued in the Borough of the Bronx, which included a larger annexation from Westchester County in 1898. The use of the definite article is attributed to the style of referring to rivers. Another explanation for the use of the definite article in the borough's name is that the original form of the name was a possessive or collective one referring to the family, as in visiting The Broncks, The Bronck's or The Broncks'.

Before 1914
The development of the Bronx is directly connected to its strategic location between New England and New York (Manhattan). Control over the bridges across the Harlem River plagued the period of British colonial rule. Kingsbridge, built in 1693 where Broadway reached the Spuyten Duyvil Creek, was a possession of the lords of Philipse Manor. The tolls they charged were resented by Bronx farmers with crops and cattle to sell in New York. It was the angry farmers who built a "free bridge" across the Harlem River which led to the abandonment of tolls altogether.

The territory now contained within Bronx County was originally part of Westchester County, one of the 12 original counties of the English Province of New York. The present Bronx County was contained in the town of Westchester and parts of the towns of Yonkers, Eastchester, and Pelham. In 1846, a new town, West Farms, was created by division of Westchester; in turn, in 1855, the town of Morrisania was created from West Farms. In 1873, the town of Kingsbridge (roughly corresponding to the modern Bronx neighborhoods of Kingsbridge, Riverdale, and Woodlawn) was established within the former borders of Yonkers.

Among famous settlers in the Bronx in the 19th and early 20th centuries were the author Willa Cather, the tobacco merchant Pierre Lorillard, and the inventor Jordan L. Mott, who established Mott Haven to house the workers at his iron works.

The consolidation of the Bronx into New York City proceeded in two stages. In 1873, the state legislature annexed Kingsbridge, West Farms and Morrisania to New York, effective in 1874; the three towns were abolished in the process. In 1895, three years before New York's consolidation with Brooklyn, Queens and Staten Island, the whole of the territory east of the Bronx River, including the Town of Westchester (which had voted in 1894 against consolidation) and portions of Eastchester and Pelham, were annexed to the city. City Island, a nautical community, voted to join the city in 1896.

On January 1, 1898, the consolidated City of New York was born, including the Bronx as one of the five distinct Boroughs. (At the same time the Bronx's territory moved from Westchester County into New York County, which already contained Manhattan and the rest of pre-1874 New York City.)

On April 19, 1912, those parts of New York County which had been annexed from Westchester County in the past decades were newly constituted as Bronx County, the 62nd and last county to be created by the state, effective in 1914. Bronx County's courts opened for business on January 2, 1914 (the same day that John P. Mitchel started work as Mayor of New York City).

Since 1914
At the end of World War I, the Bronx hosted the rather small 1918 World's Fair at 177th Street and DeVoe Avenue.

The Bronx underwent rapid growth after World War I. Extensions of the New York City Subway contributed to the increase in population as thousands of immigrants flooded the Bronx, resulting in a major boom in residential construction. Among these groups, many Irish-Americans, Italian-Americans and especially Jewish-Americans settled here. In addition, French, German, and Polish immigrants moved into the borough. The Jewish population also increased notably during this time. In 1937, according to Jewish organizations, 592,185 Jews lived in the Bronx (43.9% of the borough's population), while in 2002, only about 45,000 did. Many synagogues still stand in the Bronx, but most have been converted to other uses.

In Prohibition days (1920-33), bootleggers and gangs ran rampant in the Bronx. Irish, Italian and Polish immigrants smuggled in most of the illegal whiskey. By 1926, the Bronx was noted for its high crime rate and its many speakeasies.

After the 1930s, the Irish immigrant population in the Bronx decreased. The German population followed suit in the 1940s, as did many Italians in the 1950s and Jews in the 1960s. As the generation of the 1930s retired, many moved to southeastern Florida, west of Fort Lauderdale and Palm Beach. The migration has left a Hispanic (mostly Puerto Rican and Dominican) and African-American population, along with some non-Hispanic white areas in the far southeastern and northwestern parts of the county.

During the mid-1960s to the mid-1970s, the Bronx went into an era of sharp decline in the residents' quality of life. Historians and social scientists have put forward many factors. They include the theory (elaborated in Robert Caro's biography The Power Broker) that Robert Moses' Cross-Bronx Expressway destroyed existing residential neighborhoods. Another (unintended) factor in the Bronx's decline may have been the development of high-rise housing projects. Yet another may have been a reduction in the real-estate listings and property-related financial services (such as mortgages or insurance policies) offered in some areas of the Bronx — a process known as redlining. Others have suggested a "planned shrinkage" of municipal services, such as fire-fighting. There was also much debate as to whether rent control laws had made it less profitable (or more costly) for landlords to maintain existing buildings with their existing tenants than to abandon or destroy those buildings.

In the 1970s, the Bronx was plagued by a wave of arson. The burning of buildings was mostly in the South Bronx, concentrated especially along Westchester Avenue and in West Farms. The most common explanation of what occurred was that landlords decided to burn their buildings and take the insurance money as profit. Competing explanations blamed the insurance companies —since their non-renewals of policies might have encouraged the landlords— or the residents themselves. After the destruction of many buildings in the South Bronx, the arsons slowed significantly in the later part of the decade, but the after-effects were still felt into the early 1990s.

Since the mid-1980s, significant residential development has occurred, stimulated by the city's "Ten-Year Housing Plan" and community members working to rebuild the social, economic and environmental infrastructure by creating affordable housing. Groups affiliated with South Bronx churches erected the Nehemiah Homes with about 1,000 units. The grass roots organization Nos Quedamos' endeavor known as Melrose Commons  began to rebuild the South Bronx. The ripple effects have been felt borough-wide. As a result of the growing population, the IRT White Plains Road Line has had an increase in riders. Chains such as Marshalls, Staples and Target have opened stores in the Bronx. More bank branches have opened in the Bronx as a whole (rising from 106 in 1997 to 149 in 2007), although not primarily in poor or minority neighborhoods, while the Bronx still has fewer branches per person than wealthier boroughs.

In 1997, the Bronx was designated an All America City by the National Civic League, signifying its comeback from the decline of the 1970s. In 2006, The New York Times reported that "construction cranes have become the borough's new visual metaphor, replacing the window decals of the 1980s in which pictures of potted plants and drawn curtains were placed in the windows of abandoned buildings." The borough has experienced substantial new building construction since 2002. Between 2002 and June 2007, 33,687 new units of housing were built or were under way and $4.8 billion has been invested in new housing. In the first six months of 2007 alone total investment in new residential development was $965 million and 5,187 residential units were scheduled to be completed. Much of the new development is springing up in formerly vacant lots across the South Bronx.