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Apple From Wikipedia, the free encyclopedia Jump to navigationJump to search This article is about the fruit and the tree. For the technology company, see Apple Inc. For other uses, see Apple (disambiguation), Apple Blossom (disambiguation), and Apple tree (disambiguation). Apple Honeycrisp.jpg Fruit Malus domestica a1.jpg Flowers Scientific classification edit Kingdom:	Plantae Clade:	Angiosperms Clade:	Eudicots Clade:	Rosids Order:	Rosales Family:	Rosaceae Genus:	Malus Species:	M. pumila Binomial name Malus pumila Miller, 1768 Synonyms[1][2] Malus communis Desf. Malus domestica Borkh. nom. illeg. M. frutescens Medik. M. paradisiaca (L.) Medikus M. sylvestris Mil. Pyrus malus L. Pyrus malus var. paradisiaca L. Pyrus dioica Moench An apple is a sweet, edible fruit produced by an apple tree (Malus pumila). Apple trees are cultivated worldwide and are the most widely grown species in the genus Malus. The tree originated in Central Asia, where its wild ancestor, Malus sieversii, is still found today. Apples have been grown for thousands of years in Asia and Europe and were brought to North America by European colonists. Apples have religious and mythological significance in many cultures, including Norse, Greek and European Christian traditions.

Apple trees are large if grown from seed. Generally, apple cultivars are propagated by grafting onto rootstocks, which control the size of the resulting tree. There are more than 7,500 known cultivars of apples, resulting in a range of desired characteristics. Different cultivars are bred for various tastes and use, including cooking, eating raw and cider production. Trees and fruit are prone to a number of fungal, bacterial and pest problems, which can be controlled by a number of organic and non-organic means. In 2010, the fruit's genome was sequenced as part of research on disease control and selective breeding in apple production.

Worldwide production of apples in 2017 was 83.1 million tonnes, with China accounting for half of the total.[3]

Contents 1	Botanical information 1.1	Wild ancestors 1.2	Genome 2	History 3	Society and culture 3.1	Germanic paganism 3.2	Greek mythology 3.3	Christian art 4	Cultivars 5	Cultivation 5.1	Breeding 5.2	Pollination 5.3	Maturation and harvest 5.4	Storage 5.5	Pests and diseases 6	Production 7	Nutrition 8	Human consumption 8.1	Popular uses 8.2	Organic production 8.3	Phytochemicals 8.4	Other products 8.5	Health effects 8.6	Allergy 8.7	Toxicity of seeds 9	Proverbs 10	See also 11	References 12	Further reading 13	External links Botanical information

Blossoms, fruits, and leaves of the apple tree (Malus pumila)

An apple's side, stem end, and interior The apple is a deciduous tree, generally standing 6 to 15 ft (1.8 to 4.6 m) tall in cultivation and up to 30 ft (9.1 m) in the wild. When cultivated, the size, shape and branch density are determined by rootstock selection and trimming method. The leaves are alternately arranged dark green-colored simple ovals with serrated margins and slightly downy undersides.[4]

Apple blossom Blossoms are produced in spring simultaneously with the budding of the leaves and are produced on spurs and some long shoots. The 3 to 4 cm (1.2 to 1.6 in) flowers are white with a pink tinge that gradually fades, five petaled, with an inflorescence consisting of a cyme with 4–6 flowers. The central flower of the inflorescence is called the "king bloom"; it opens first and can develop a larger fruit.[4][5]

The fruit matures in late summer or autumn, and cultivars exist in a wide range of sizes. Commercial growers aim to produce an apple that is 2 3⁄4 to 3 1⁄4 in (7.0 to 8.3 cm) in diameter, due to market preference. Some consumers, especially those in Japan, prefer a larger apple, while apples below 2 1⁄4 in (5.7 cm) are generally used for making juice and have little fresh market value. The skin of ripe apples is generally red, yellow, green, pink, or russetted, though many bi- or tri-colored cultivars may be found.[6] The skin may also be wholly or partly russeted i.e. rough and brown. The skin is covered in a protective layer of epicuticular wax.[7] The exocarp (flesh) is generally pale yellowish-white,[6] though pink or yellow exocarps also occur.

Wild ancestors Main article: Malus sieversii The original wild ancestor of Malus pumila was Malus sieversii, found growing wild in the mountains of Central Asia in southern Kazakhstan, Kyrgyzstan, Tajikistan, and Xinjiang, China.[4][8] Cultivation of the species, most likely beginning on the forested flanks of the Tian Shan mountains, progressed over a long period of time and permitted secondary introgression of genes from other species into the open-pollinated seeds. Significant exchange with Malus sylvestris, the crabapple, resulted in current populations of apples being more related to crabapples than to the more morphologically similar progenitor Malus sieversii. In strains without recent admixture the contribution of the latter predominates.[9][10][11]

Genome In 2010, an Italian-led consortium announced they had sequenced the complete genome of the apple in collaboration with horticultural genomicists at Washington State University,[12] using 'Golden Delicious'.[13] It had about 57,000 genes, the highest number of any plant genome studied to date[14] and more genes than the human genome (about 30,000).[15] This new understanding of the apple genome will help scientists identify genes and gene variants that contribute to resistance to disease and drought, and other desirable characteristics. Understanding the genes behind these characteristics will help scientists perform more knowledgeable selective breeding. The genome sequence also provided proof that Malus sieversii was the wild ancestor of the domestic apple—an issue that had been long-debated in the scientific community.[12]

History color photograph of a hand holding a red apple Wild Malus sieversii apple in Kazakhstan The center of diversity of the genus Malus is in eastern present-day Turkey. The apple tree may have been the earliest tree that humans cultivated,[16] and growers have improved its fruits through selection over thousands of years. Alexander the Great is credited with finding dwarfed apples in Kazakhstan in 328 BCE.[4] Winter apples, picked in late autumn and stored just above freezing, have been an important food in Asia and Europe for millennia.[16]

Of the many Old World plants that the Spanish introduced to Chiloé Archipelago in the 16th century, apple trees became particularly well adapted.[17] Apples were introduced to North America by colonists in the 17th century,[4] and the first apple orchard on the North American continent was planted in Boston by Reverend William Blaxton in 1625.[18] The only apples native to North America are crab apples, which were once called "common apples".[19] Apple cultivars brought as seed from Europe were spread along Native American trade routes, as well as being cultivated on colonial farms. An 1845 United States apples nursery catalogue sold 350 of the "best" cultivars, showing the proliferation of new North American cultivars by the early 19th century.[19] In the 20th century, irrigation projects in Eastern Washington began and allowed the development of the multibillion-dollar fruit industry, of which the apple is the leading product.[4]

Until the 20th century, farmers stored apples in frostproof cellars during the winter for their own use or for sale. Improved transportation of fresh apples by train and road replaced the necessity for storage.[20][21] Controlled atmosphere facilities are used to keep apples fresh year-round. Controlled atmosphere facilities use high humidity, low oxygen, and controlled carbon dioxide levels to maintain fruit freshness. They were first used in the United States in the 1960s.[22]

Society and culture Main article: Apple (symbolism) Germanic paganism Illustration of girl in a red dress, holding 3 candles in one hand and a basket of apples in the other "Brita as Iduna" (1901) by Carl Larsson In Norse mythology, the goddess Iðunn is portrayed in the Prose Edda (written in the 13th century by Snorri Sturluson) as providing apples to the gods that give them eternal youthfulness. English scholar H. R. Ellis Davidson links apples to religious practices in Germanic paganism, from which Norse paganism developed. She points out that buckets of apples were found in the Oseberg ship burial site in Norway, that fruit and nuts (Iðunn having been described as being transformed into a nut in Skáldskaparmál) have been found in the early graves of the Germanic peoples in England and elsewhere on the continent of Europe, which may have had a symbolic meaning, and that nuts are still a recognized symbol of fertility in southwest England.[23]

Davidson notes a connection between apples and the Vanir, a tribe of gods associated with fertility in Norse mythology, citing an instance of eleven "golden apples" being given to woo the beautiful Gerðr by Skírnir, who was acting as messenger for the major Vanir god Freyr in stanzas 19 and 20 of Skírnismál. Davidson also notes a further connection between fertility and apples in Norse mythology in chapter 2 of the Völsunga saga when the major goddess Frigg sends King Rerir an apple after he prays to Odin for a child, Frigg's messenger (in the guise of a crow) drops the apple in his lap as he sits atop a mound.[24] Rerir's wife's consumption of the apple results in a six-year pregnancy and the Caesarean section birth of their son—the hero Völsung.[25]

Further, Davidson points out the "strange" phrase "Apples of Hel" used in an 11th-century poem by the skald Thorbiorn Brúnarson. She states this may imply that the apple was thought of by Brúnarson as the food of the dead. Further, Davidson notes that the potentially Germanic goddess Nehalennia is sometimes depicted with apples and that parallels exist in early Irish stories. Davidson asserts that while cultivation of the apple in Northern Europe extends back to at least the time of the Roman Empire and came to Europe from the Near East, the native varieties of apple trees growing in Northern Europe are small and bitter. Davidson concludes that in the figure of Iðunn "we must have a dim reflection of an old symbol: that of the guardian goddess of the life-giving fruit of the other world."[23]

Greek mythology

Heracles with the apple of Hesperides Apples appear in many religious traditions, often as a mystical or forbidden fruit. One of the problems identifying apples in religion, mythology and folktales is that the word "apple" was used as a generic term for all (foreign) fruit, other than berries, including nuts, as late as the 17th century.[26] For instance, in Greek mythology, the Greek hero Heracles, as a part of his Twelve Labours, was required to travel to the Garden of the Hesperides and pick the golden apples off the Tree of Life growing at its center.[27][28][29]

The Greek goddess of discord, Eris, became disgruntled after she was excluded from the wedding of Peleus and Thetis.[30] In retaliation, she tossed a golden apple inscribed Καλλίστη (Kalliste, sometimes transliterated Kallisti, "For the most beautiful one"), into the wedding party. Three goddesses claimed the apple: Hera, Athena, and Aphrodite. Paris of Troy was appointed to select the recipient. After being bribed by both Hera and Athena, Aphrodite tempted him with the most beautiful woman in the world, Helen of Sparta. He awarded the apple to Aphrodite, thus indirectly causing the Trojan War.[31]

The apple was thus considered, in ancient Greece, sacred to Aphrodite. To throw an apple at someone was to symbolically declare one's love; and similarly, to catch it was to symbolically show one's acceptance of that love. An epigram claiming authorship by Plato states:[32]

I throw the apple at you, and if you are willing to love me, take it and share your girlhood with me; but if your thoughts are what I pray they are not, even then take it, and consider how short-lived is beauty.

— Plato, Epigram VII Atalanta, also of Greek mythology, raced all her suitors in an attempt to avoid marriage. She outran all but Hippomenes (also known as Melanion, a name possibly derived from melon the Greek word for both "apple" and fruit in general),[28] who defeated her by cunning, not speed. Hippomenes knew that he could not win in a fair race, so he used three golden apples (gifts of Aphrodite, the goddess of love) to distract Atalanta. It took all three apples and all of his speed, but Hippomenes was finally successful, winning the race and Atalanta's hand.[27]

Christian art

Adam and Eve by Albrecht Dürer (1507), showcasing the apple as a symbol of sin. Though the forbidden fruit of Eden in the Book of Genesis is not identified, popular Christian tradition has held that it was an apple that Eve coaxed Adam to share with her.[33] The origin of the popular identification with a fruit unknown in the Middle East in biblical times is found in confusion between the Latin words mālum (an apple) and mălum (an evil), each of which is normally written malum.[34] The tree of the forbidden fruit is called "the tree of the knowledge of good and evil" in Genesis 2:17, and the Latin for "good and evil" is bonum et malum.[35]

Renaissance painters may also have been influenced by the story of the golden apples in the Garden of Hesperides. As a result, in the story of Adam and Eve, the apple became a symbol for knowledge, immortality, temptation, the fall of man into sin, and sin itself. The larynx in the human throat has been called the "Adam's apple" because of a notion that it was caused by the forbidden fruit remaining in the throat of Adam.[33] The apple as symbol of sexual seduction has been used to imply human sexuality, possibly in an ironic vein.[33]

Cultivars Main article: List of apple cultivars There are more than 7,500 known cultivars of apples.[36] Cultivars vary in their yield and the ultimate size of the tree, even when grown on the same rootstock.[37] Different cultivars are available for temperate and subtropical climates. The UK's National Fruit Collection, which is the responsibility of the Department of Environment, Food, and Rural Affairs, includes a collection of over 2,000 cultivars of apple tree in Kent.[38] The University of Reading, which is responsible for developing the UK national collection database, provides access to search the national collection. The University of Reading's work is part of the European Cooperative Programme for Plant Genetic Resources of which there are 38 countries participating in the Malus/Pyrus work group.[39]

The UK's national fruit collection database contains a wealth of information on the characteristics and origin of many apples, including alternative names for what is essentially the same "genetic" apple cultivar. Most of these cultivars are bred for eating fresh (dessert apples), though some are cultivated specifically for cooking (cooking apples) or producing cider. Cider apples are typically too tart and astringent to eat fresh, but they give the beverage a rich flavor that dessert apples cannot.[40]

Commercially popular apple cultivars are soft but crisp. Other desirable qualities in modern commercial apple breeding are a colorful skin, absence of russeting, ease of shipping, lengthy storage ability, high yields, disease resistance, common apple shape, and developed flavor.[37] Modern apples are generally sweeter than older cultivars, as popular tastes in apples have varied over time. Most North Americans and Europeans favor sweet, subacid apples, but tart apples have a strong minority following.[41] Extremely sweet apples with barely any acid flavor are popular in Asia,[41] especially the Indian Subcontinent .[40]

Old cultivars are often oddly shaped, russeted, and grow in a variety of textures and colors. Some find them to have better flavor than modern cultivars,[42] but they may have other problems that make them commercially unviable—low yield, disease susceptibility, poor tolerance for storage or transport, or just being the "wrong" size. A few old cultivars are still produced on a large scale, but many have been preserved by home gardeners and farmers that sell directly to local markets. Many unusual and locally important cultivars with their own unique taste and appearance exist; apple conservation campaigns have sprung up around the world to preserve such local cultivars from extinction. In the United Kingdom, old cultivars such as 'Cox's Orange Pippin' and 'Egremont Russet' are still commercially important even though by modern standards they are low yielding and susceptible to disease.[4]

'Alice'

'Ambrosia'

'Ananasrenette'

'Arkansas Black'

'Aroma'

'Belle de Boskoop'

'Bramley'

'Cox Pomona'

'Discovery'

'Fuji'

'Gala'

'Golden Delicious'

'Goldrenette', ('Reinette')

'Granny Smith'

'Honeycrisp'

'James Grieve'

'Jonagold'

'Lobo'

'McIntosh'

'Pacific rose'

'Pink Lady'

'Red Delicious'

'Sampion' (Shampion)

'Stark Delicious' (ripening)

'SugarBee'

'Summerred'

'Yellow Transparent'