User:Rupert Clayton/Draft of List of Sisyrinchium species

Approximately 211 species are currently accepted within the genus Sisyrinchium. Botanists have divided the genus using several schemes in the past, but the current division into 10 sections is rooted in phylogenetic analysis and mapped to consistent morphological characters.

This list shows currently accepted species, subspecies and varieties of Sisyrinchium for each section. Unless otherwise stated, this article follows Plants of the World Online (POWO) for accepted taxa and synonyms. The foundation for placement of species into sections is a 2017 study by Camila Dellanhese Inácio and collaborators that analyzed 171 samples from 110 Sisyrinchium taxa (96 published species, 3 infrataxa, and 11 unpublished taxa from south and southeast Brazil). That study identified 10 distinct clades within the genus, into which 108 of the 110 taxa could be placed with confidence. Two species, S. chilense and S. elegantulum, showed conclusive indications of reticulate evolution, with parentage from more than one section.

Beyond the assignments explicitly made by Inácio et al, there are several reasons that other species can be confidently assigned to one of the 10 sections. Some sections are practically identical to a section or subgenus proposed by previous writers. An example is Inácio's Section Sisyrinchium, which corresponds to Section Bermudiana sensu Benth. & Hook and includes all the North American species placed in Section Sisyrinchium by Pierfelice Ravenna. There are also some scenarios where Inácio's paper provides sufficient information to confidently assign taxa to sections, such as assigning an infrataxon to the same section as its parent or sibling taxon. Section assignments can also be made for undescribed taxa studied by Inácio that have since been formally described. And several papers describing new taxa since 2017 have used Inácio's terminology to determine the appropriate section for the new species.

Section Hydastylus
Phylogenetic analysis by Inácio et al identified Section Hydastylus as the first lineage to diverge from the genus, and thus as a sister to all the other sections. Morphologically, plants in this section can be distinguished by the combination of having inflorescences with a single terminal rhipidium, and basal leaves but no cauline leaves. Species in Section Hydastylus are similiar to those in Section Segetia in many respects, but lack any bract below the spathes of the inflorescence and have stems that are obviously winged. They have yellow, disk-shaped flowers without trichomes (in common with many other sections). Species within Section Hydastylus are found across a wide area from the Páramo and Puna regions of the northern Andes, through Central America and the mountains of Mexico to western North America as far north as Vancouver Island. One species, S. acre, grows in Hawaii.

• Sisyrinchium acre H.Mann

• Sisyrinchium arguellesiae Ceja, Espejo & López-Ferr.

• Sisyrinchium brachypus (E.P.Bicknell) J.K.Henry

• Sisyrinchium californicum (Ker-Gawl.) Dryand. – golden blue-eyed grass, yellow-eyed-grass

• Sisyrinchium cernuum (Bickn.) Kearney – nodding blue-eyed grass

• Sisyrinchium elmeri Greene

• Sisyrinchium longipes (Bickn.) Kearney & Peebles – timberland blue-eyed grass

• Sisyrinchium palustre Diels

• Sisyrinchium schaffneri S.Wats.

• Sisyrinchium serrulatum (E.P.Bicknell) Espejo & López-Ferr.

• Sisyrinchium subalpinum Henrich & Goldblatt

• Sisyrinchium subcernuum (E.P.Bicknell) Henrich & Goldblatt

• Sisyrinchium tinctorium Kunth

• Sisyrinchium translucens (E.P.Bicknell) Espejo & López-Ferr.


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Section Segetia
• Sisyrinchium brevipes Baker

• Sisyrinchium caespitificum Kraenzl.

• Sisyrinchium humahuacense C.A. Zanotti

• Sisyrinchium jamesonii Baker

• Sisyrinchium laxinervium Ravenna

• Sisyrinchium praealtum Kranzl.

• Sisyrinchium pusillum Kunth

• Sisyrinchium trinerve Baker


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Section Echthronema
• Sisyrinchium angustissimum (B.L.Rob. & Greenman) Greenman & C.Thompson

• Sisyrinchium arizonicum Rothrock – Arizona blue-eyed grass

• Sisyrinchium chiricanum Woodson

• Sisyrinchium convolutum Nocca

• Sisyrinchium macrophyllum Greenman

• Sisyrinchium mandonii Baker

• Sisyrinchium tenuifolium Humb. & Bonpl. ex Willd.


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Section Spathirachis
• Sisyrinchium arenarium Poeppig subsp. arenarium – yellow-eyed grass

• Sisyrinchium arenarium subsp. adenostemon (Phil.) Ravenna

• Sisyrinchium chapelcoense Ravenna

• Sisyrinchium convallium Ravenna

• Sisyrinchium cuspidatum Poeppig

• Sisyrinchium graminifolium Lindley

• Sisyrinchium limarinum Ravenna

• Sisyrinchium macrocarpum Hieron.

• Sisyrinchium papillosum R.C.Foster

• Sisyrinchium pearcei Phil.

• Sisyrinchium striatum Sm. – pale yellow-eyed-grass or satin flower


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Section Viperella
• Sisyrinchium alatum Hook.

• Sisyrinchium angustius (Ravenna) C.D.Inácio & L.Eggers

• Sisyrinchium balansae Baker

• Sisyrinchium brasiliense (Ravenna) Ravenna

• Sisyrinchium bromelioides R.C.Foster

• Sisyrinchium burchellii Baker

• Sisyrinchium caeteanum Ravenna

• Sisyrinchium coalitum Ravenna

• Sisyrinchium congestum Klatt

• Sisyrinchium decumbens Ravenna

• Sisyrinchium densiflorum Ravenna

• Sisyrinchium diversicarpum C.D.Inácio & L.Eggers

• Sisyrinchium flabellatum Aita & L.Eggers

• Sisyrinchium macrocephalum Graham

• Sisyrinchium marginatum Klatt

• Sisyrinchium nidulare (Hand.-Mazz.) I.M.Johnst.

• Sisyrinchium oxyspathum Ravenna

• Sisyrinchium palmifolium subsp. fuscoviride (Ravenna) Ravenna

• Sisyrinchium palmifolium L. subsp. palmifolium

• Sisyrinchium paludosum (Ravenna) C.D.Inácio & L.Eggers

• Sisyrinchium plicatulum Ravenna

• Sisyrinchium rectilineum Ravenna

• Sisyrinchium restioides Spreng.

• Sisyrinchium vaginatum subsp. ciliolatum Ravenna

• Sisyrinchium vaginatum subsp. marchio (Vell.) Ravenna

• Sisyrinchium vaginatum Spreng. subsp. vaginatum

• Sisyrinchium weirii Baker

• Sisyrinchium wettsteinii Handel-Mazzetti


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Section Cephalanthum
• Sisyrinchium albilapidense Ravenna

• Sisyrinchium claritae Herter

• Sisyrinchium commutatum subsp. capillare Ravenna

• Sisyrinchium commutatum Klatt subsp. commutatum

• Sisyrinchium commutatum subsp. piliferum (Klatt) C.D.Inácio & L.Eggers

• Sisyrinchium dasyspathum (Ravenna) Ravenna

• Sisyrinchium fasciculatum Klatt

• Sisyrinchium fiebrigii I.M.Johnst.

• Sisyrinchium hasslerianum Baker

• Sisyrinchium hoehnei I.M.Johnst.

• Sisyrinchium luzula Klotzsch ex Klatt

• Sisyrinchium ostenianum Beauverd

• Sisyrinchium pampeanum C.D.Inácio & L.Eggers

• Sisyrinchium pendulum (Ravenna) Ravenna

• Sisyrinchium platyphyllum S. Wats.

• Sisyrinchium purpurellum Ravenna subsp. purpurellum

• Sisyrinchium purpurellum subsp. trichospathum Ravenna

• Sisyrinchium rambonis R.C.Foster

• Sisyrinchium scariosum I.M.Johnst

• Sisyrinchium sectiandrum C.D.Inácio & L.Eggers

• Sisyrinchium sellowianum Klatt

• Sisyrinchium setaceum Klatt

• Sisyrinchium soboliferum Ravenna

• Sisyrinchium subnudum I.M.Johnst.

• Sisyrinchium teleanthum Ravenna

• Sisyrinchium trichospathum Ravenna


 * Sources

Section Trichoparcus
• Sisyrinchium avenaceum Klatt

• Sisyrinchium foliosum I.M.Johnst

• Sisyrinchium megapotamicum Malme

• Sisyrinchium rosengurttii I.M.Johnst.

• Sisyrinchium uliginosum Ravenna

• Sisyrinchium valparadiseum Ravenna


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Section Morphanthus
• Sisyrinchium laxum Otto – veined yellow-eyed-grass

• Sisyrinchium micranthum Cav. – blue pigroot or striped rush-leaf

• Sisyrinchium rosulatum Bickn. – annual blue-eyed grass


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Section Rhizilineum
• Sisyrinchium minus Engelm. & A.Gray – dwarf blue-eyed grass

• Sisyrinchium minutiflorum Klatt


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Section Sisyrinchium
• Sisyrinchium albidum Raf. – white blue-eyed grass

• Sisyrinchium angustifolium Mill – narrow-leaf blue-eyed-grass

• Sisyrinchium atlanticum Bickn. – eastern blue-eyed grass

• Sisyrinchium azureum Phil.

• Sisyrinchium bellum S.Watson – western blue-eyed grass

• Sisyrinchium bermudiana  – blue-eyed grass

• Sisyrinchium biforme Bickn. – wiry blue-eyed grass

• Sisyrinchium calciphilum Sorrie

• Sisyrinchium campestre Bickn. – prairie blue-eyed grass

• Sisyrinchium capillare Bickn.

• Sisyrinchium demissum Greene – stiff blue-eyed grass

• Sisyrinchium dichotomum Bickn. – wishbone blue-eyed grass

• Sisyrinchium elizabethiae E.L.Keith

• Sisyrinchium ensigerum E.P.Bicknell

• Sisyrinchium funereum Bickn. – Funeral Mountain blue-eyed grass

• Sisyrinchium fuscatum Bickn. – coastalplain blue-eyed grass

• Sisyrinchium groenlandicum Böcher

• Sisyrinchium halophilum Greene – Nevada blue-eyed grass

• Sisyrinchium hitchcockii D.Henderson – Hitchcock's blue-eyed grass

• Sisyrinchium idahoense Bickn. var. idahoense – Idaho blue-eyed grass

• Sisyrinchium idahoense var. macounii (E.P.Bicknell) Douglass M.Hend.

• Sisyrinchium idahoense var. occidentale (E.P.Bicknell) Douglass M.Hend.

• Sisyrinchium idahoense var. segetum (E.P.Bicknell) Douglass M.Hend.

• Sisyrinchium langloisii Greene

• Sisyrinchium littorale Greene – Alaska blue-eyed grass

• Sisyrinchium macranthum Griseb.

• Sisyrinchium miamiense Bickn. – Miami blue-eyed grass

• Sisyrinchium montanum Greene – strict blue-eyed grass, American blue-eyed grass

• Sisyrinchium mucronatum Michx. – needletip blue-eyed grass

• Sisyrinchium myrioflorum E.L.Keith

• Sisyrinchium nashii Bickn. – Nash's blue-eyed grass

• Sisyrinchium novoleonense G.L.Nesom & L.Hern.

• Sisyrinchium pachyrhizum subsp. columellosum Ravenna

• Sisyrinchium pachyrhizum Baker subsp. pachyrhizum

• Sisyrinchium pachyrhizum subsp. procerum Ravenna

• Sisyrinchium pallidum Cholewa & D.Henderson – pale blue-eyed grass

• Sisyrinchium patagonicum Phil. ex Baker

• Sisyrinchium platense I.M.Johnst

• Sisyrinchium pruinosum  – spotted blue-eyed grass

• Sisyrinchium radicatum Bickn. – bigroot blue-eyed grass

• Sisyrinchium sagittiferum Bickn. – spearbract blue-eyed grass

• Sisyrinchium sarmentosum Suksdorf ex Greene – mountain blue-eyed grass

• Sisyrinchium scabrum Schltdl. & Cham.

• Sisyrinchium septentrionale Bickn. – northern blue-eyed grass

• Sisyrinchium strictum Bickn.

• Sisyrinchium xerophyllum Greene – jeweled blue-eyed grass


 * Sources

Species unplaced due to hybrid origin
• Sisyrinchium chilense Hook – swordleaf blue-eyed grass

• Sisyrinchium elegantulum Ravenna


 * Sources

Species yet to be placed
The following species are accepted by Plants of the World Online but have not been placed in a section following the taxonomy of Inácio et al 2017. • Sisyrinchium abietum McVaugh

• Sisyrinchium anadenicum Ravenna

• Sisyrinchium andinopatagonicum Ravenna

• Sisyrinchium annuum Ravenna

• Sisyrinchium antucense Ravenna

• Sisyrinchium aquidaunicum Ravenna

• Sisyrinchium arrayanicum Ravenna

• Sisyrinchium biflorum Griseb.

• Sisyrinchium binervatum Ravenna

• Sisyrinchium breviarmium Ravenna

• Sisyrinchium calathinum Ravenna

• Sisyrinchium chaguaranicum Ravenna

• Sisyrinchium cholewae Espejo, López-Ferr. & Ceja

• Sisyrinchium conzattii Calderón & Rzed.

• Sisyrinchium coulterianum Klatt ex Baker

• Sisyrinchium cylindrocarpium Herter

• Sisyrinchium cyperellum Ravenna

• Sisyrinchium deflexum R.C.Foster

• Sisyrinchium deseadense Ravenna

• Sisyrinchium eserrulatum I.M.Johnst.

• Sisyrinchium exalatum B.L.Rob. & Greenman

• Sisyrinchium exilifolium Ravenna

• Sisyrinchium firmifolium Ravenna

• Sisyrinchium galapagense Ravenna

• Sisyrinchium glaziovii Baker

• Sisyrinchium gratissimum Ravenna

• Sisyrinchium guanajuatense Ceja, Espejo & López-Ferr

• Sisyrinchium hintoniorum G.L.Nesom

• Sisyrinchium hirsutum Baker

• Sisyrinchium humidum Ravenna

• Sisyrinchium hypsophilum I.M.Johnst. ex R.C.Foster

• Sisyrinchium igatimiense Ravenna

• Sisyrinchium inclinatum Ravenna

• Sisyrinchium intihuatanense (Vargas) Ravenna

• Sisyrinchium itabiritense Ravenna

• Sisyrinchium ivanii R.C.Foster

• Sisyrinchium johnstonii Standl.

• Sisyrinchium laetum (Ravenna) J.M.Watson & A.R.Flores

• Sisyrinchium laevigatum Ravenna

• Sisyrinchium laterale Baker

• Sisyrinchium latiusculum Ravenna

• Sisyrinchium longispathum Conz.

• Sisyrinchium magnicapsulare J.M.Watson

• Sisyrinchium maipoanum Ravenna

• Sisyrinchium marchioides Ravenna

• Sisyrinchium martense Ravenna

• Sisyrinchium microbracteatum G.L.Nesom

• Sisyrinchium nanum Phil.

• Sisyrinchium nembyense Ravenna

• Sisyrinchium nigricans Gay

• Sisyrinchium obconicum Ravenna

• Sisyrinchium orbiculatum Ravenna

• Sisyrinchium paramorum Ravenna

• Sisyrinchium parvifolium Baker

• Sisyrinchium paucispathum Ravenna

• Sisyrinchium planicola Ceja & Cholewa

• Sisyrinchium pringlei B.L.Rob. & Greenman

• Sisyrinchium quadrangulatum Klatt

• Sisyrinchium rectivalvatum Ravenna

• Sisyrinchium reitzii R.C.Foster

• Sisyrinchium rigidifolium Baker

• Sisyrinchium scalarium Ravenna

• Sisyrinchium schottii Herter

• Sisyrinchium sinuosum Ravenna

• Sisyrinchium somuncurense Ravenna

• Sisyrinchium subtile Ravenna

• Sisyrinchium tafiense Ravenna

• Sisyrinchium tofoense Ravenna

• Sisyrinchium tolucense Peyr.

• Sisyrinchium trichanthum Dusén

• Sisyrinchium tucumanum Ravenna

• Sisyrinchium unguiculatum Griseb.

• Sisyrinchium unispathaceum Klatt

• Sisyrinchium venezolense Ravenna

• Sisyrinchium vestitum Ravenna

• Sisyrinchium zamudioi Espejo, López-Ferr. & Ceja

Formerly placed here

 * Calydorea speciosa (Hook.) Herb. (as S. speciosum Hook. )
 * Eleutherine bulbosa (Mill.) Urb. (as S. bulbosum Mill. )
 * Libertia ixioides (G.Forst.) Spreng. (as S. ixioides G.Forst. )
 * Solenomelus pedunculatus (Gillies) Hochr. (as S. pedunculatum Gillies ex Hook. )
 * Olsynium junceum