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Richard F. Shaw
born May 14, 1924, Redlands, California, died May 13, 2015, Bethesda, Maryland.

Richard F. Shaw, with J.D. Mohler, was the author of an influential 1953 paper "The Selective Significance of the Sex Ratio". Known for his development of the Shaw-Mohler equation (also known as the Shaw-Mohler theorem), a method in population genetics used to determine the ratio of a gene or trait in a population over time. The method was introduced to a new generation by Eric Charnov in his book "The Theory of Sex Allocation" in 1982 and again in C.G. Williams' book "Adaptation and Natural Selection" (1996), where he calls the 1953 paper which introduced the Shaw-Mohler equation 'superb' and credits it as a major influence on his thinking. Though he is now known for the Shaw-Mohler equation, most of his professional career was focused on the study of genetic diseases such as Muscular Dystrophy, Tay-Sachs and Cycle Cell anemia. His autobiography, Itinerant Professor: A Memoir, about his early life and peripatetic academic career is available on Amazon.

Some of his papers
Shaw RF. The question of natural selection against mutable-to-terminator codons. J Hered. 1991 Jan-Feb;82(1):77-80. .

Shaw RF, Bloom RW, Bowman JE. Hemoglobin and the genetic code. Evolution of protection against somatic mutation. J Mol Evol. 1977 May 13;9(3):225-30. .

Shaw RF, Kallen RJ. Population genetics of Alport's syndrome. Hypothesis of abnormal segregation and the necessary existence of mutation. Nephron. 1976;16(6):427-32..

MacNeill E, Shaw RF. Segregation ratios in Alport's syndrome. J Med Genet. 1973 Mar;10(1):23-6. ;.

Shaw RF, Gall JC Jr, Schuman SH. Febrile convulsions as a problem in waiting times. Epilepsia. 1972 Apr;13(2):305-11. .

Shaw RF, Smith AP. Strontium-90 and infant mortality in Canada. Nature. 1970 Nov 14;228(5272):667-9. .

Shaw RF. Good genes and bad: population genetics and health. N S Med Bull. 1970 Apr;49(2):41-3. .

Shaw RF, Smith AP. Is Tay-Sachs disease increasing? Nature. 1969 Dec 20;224(5225):1214-5..

Shaw RF, Dreifuss FE. Mild and severe forms of X-linked muscular dystrophy. Arch Neurol. 1969 May;20(5):451-60. .

Shaw RF, Pearson CM, Chowdhury SR, Dreifuss FE. Serum enzymes in sex-linked (Duchenne) muscular dystrophy. Arch Neurol. 1967 Feb;16(2):115-22. .

Shaw RF. The relative role of heredity and environment in congenital heart disease. A review. J Lancet. 1965 Nov;85(11):519-24. Review. .

Shaw RF. Style Manual for Biological Journals: Approval of Manuscripts. Science. 1961 Apr 21;133(3460):1275-6. .

Shaw RF, Glover RA. Abnormal Segregation in Hereditary Renal Disease with Deafness. Am J Hum Genet. 1961 Mar;13(1 Pt 1):89-97. ;.

Shaw RF. The Theoretical Genetics of the Sex Ratio. Genetics. 1958 Mar;43(2):149-63. ;.