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Hydrocotyle phoenix is a species of annual pennywort and is commonly called Fire Pennywort. It is only known to grow in south-west Australia, specifically in fire prone habitats; it is unique in this regard as it is the only species of Hydrocotyle known to have a fire adapted life-history. The specific epiphet "phoenix" references this fire adapted life history as in Greek mythology, a Phoenix experiences rebirth by rising from ashes, much like how this plant rises from the ashes after a wildfire.

Description
H. phoenix is a low growing annual herb which grows 4-15cm tall and up to 100cm wide with a sprawling habit. The leaves grow from a basal rosette. The leaves are palmately lobed and 5-20 long by 7-26mm wide and covered in small hairs. The flowers are borne in a compact umbel with about 10-20 flowers. The flowers themselves are small, only 4-6mm wide and white. The flowers have only 5 white petals and lack a calyx. The fruits are schizocarps about 1mm in size.

Atypically for this group of plants, H. phoenix is a fire-ephemeral plant, meaning that it forms a soil seed bank in the soil and its germination is triggered by wildfire. It will typically not grow unless prompted by a wildfire. Like many plants from Australia, its life cycle requires fire.

Hydrocotyle phoenix was discovered in 2015. It is most morphologically most similar to Hydrocotyle hispidula, Hydrocotyle blepharocarpa, and Hydrocotyle geraniifolia. It's evolutionary position within Hydrocotyle has not yet been determined.

Range
H. phoenix is only known from Karri Forest within D'Entrecasteaux National Park in Western Australia. The plant is locally abundant, but due to having a very small range, it is currently listed as Priority 2 as not enough data exists to ascribe it conservation code.