Hyantis

Hyantis is a monotypic nymphalid butterfly genus. Its sole species is Hyantis hodeva, which is found in New Guinea. It is uncertain which tribe this butterfly should be placed within.

Distribution
The species is distributed in the following islands: New Guinea, Waigeo, Misool, Yapen and D'Entrecasteaux Islands.

Morphology
The species has three pairs of six well-developed ocellus on the ventral side of the wings: one pair on the forewings near the apex and two pairs on the hindwings. Each eyespot is large and bordered with yellow, and the black "pupil" has plural small white dots inside. This species is similar to Taenaris, but can be easily identified from that genus, which generally has only two or four ocellus with single white spot on its hindwings.

This species is also unique in having a closed discoidal cell on each hindwing. This morphological character is also shared by the genus Morphopsis and is generally found in the subfamily Satyrinae.

The early stage is unknown.

Mimicry
This species is very similar to Taenaris species, especially T. catops. Hyantis is also imitated by Mycalesis drusillodes (Satyrini), Taenaris by Elymnias agondas (Elymniini). It should be considered that these species may be involved in some kind of mimicry complex, but the details are not known.

Systematics
The genus Hyantis (and putatively related Morphopsis) has traditionally been placed within tribe Amathusiini, even though there are differences in wing venation structures. Recent molecular phylogenetic analyses have been gradually resolved the confusion in the phylogenetic relationships in the subfamily Satyrinae, but the phylogenetic status of this genus remains confused, with several methods of analysis showing different results. For example: within the clade of Melanitini + Dirini (#|Peña & Wahlberg 2008), appears to be related with Elymnias, as sister to Amathusiini (#|Wahlberg et al. 2009, Fig.5s) and within Zetherini (#|Wahlberg et al. 2009, Fig.3s and #|Penz 2017). In any case, further studies are needed to clarify the phylogenetic position of this genus.

Classification
This species contains 3 to 5 subspecies. The lower classification shown here is according to, which recognizes 4 subspecies.


 * Hyantis Hewitson, 1862
 * Hyantis hodeva Hewitson, 1862
 * Hyantis hodeva hodeva Hewitson, 1862
 * Hyantis hodeva helvola Stichel, 1905 (syn. Hyantis hodeva hageni Röber, 1903 )
 * Hyantis hodeva fulginosa Grose-Smith, 1898 (syn. Hyantis hodeva xanthophthalma Röber, 1903 )
 * Hyantis hodeva emarginata Fruhstorfer, 1916