Portal:Amphibians
The Amphibian Portal
Amphibians are ectothermic, anamniotic, four-limbed vertebrate animals that constitute the class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all tetrapods, excluding the amniotes (tetrapods with an amniotic membrane, such as modern reptiles, birds, and mammals). All extant (living) amphibians belong to the monophyletic subclass Lissamphibia, with three living orders: Anura (frogs), Urodela (salamanders), and Gymnophiona (caecilians). Evolved to be mostly semiaquatic, amphibians have adapted to inhabit a wide variety of habitats, with most species living in freshwater, wetland or terrestrial ecosystems (such as riparian woodland, fossorial and even arboreal habitats). Their life cycle typically starts out as aquatic larvae with gills known as tadpoles, but some species have developed behavioural adaptations to bypass this.
The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards, but unlike reptiles and other amniotes, require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in recent decades there has been a dramatic decline in amphibian populations for many species around the globe.
The earliest amphibians evolved in the Devonian period from sarcopterygian fish with lungs and bony-limbed fins, features that were helpful in adapting to dry land. They diversified and became ecologically dominant during the Carboniferous and Permian periods, but were later displaced in terrestrial environments by early reptiles and basal synapsids (mammal predecessors). The origin of modern amphibians belonging to Lissamphibia, which first appeared during the Early Triassic, around 250 million years ago, has long been contentious. However the emerging consensus is that they likely originated from temnospondyls, the most diverse group of prehistoric amphibians, during the Permian period. (Full article...)
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Selected salamander article
The Salamandroidea are a suborder of salamanders, referred to as advanced salamanders. The members of the suborder are found worldwide except for Antarctica, sub-Saharan Africa, and Oceania. They differ from suborder Cryptobranchoidea as the angular and prearticular bones in their lower jaws are fused, their trunk ribs are bicapitate, and all members use internal fertilization. The female is fertilized by means of a spermatophore, a sperm-containing cap placed by the male in her cloaca. The sperm is stored in spermathecae on the roof of the cloaca until it is needed at the time of oviposition.
The earliest known salamandroid fossils remain contested. Some studies suggest that the earliest salamandroids are represented by specimens of the species Beiyanerpeton jianpingensis and Qinglongtriton gangouensis from the Tiaojishan Formation of Inner Mongolia, China, dated to the Late Jurassic about 157 million years ago. Alternative analyses suggest that Beiyanerpeton jianpingensis and Qinglongtriton gangouensis are stem salamanders and that the oldest known certain salamandroid is Valdotriton gracilis from the Early Cretaceous of Spain, about 127 Ma. (Full article...)List of selected salamander articles
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Did you know? –
- ... that the larvae of the terrestrial Ceylon caecilian have gills and a tail fin and develop in water?
- ... that the Georgia blind salamander and the Dougherty Plain cave crayfish both live underground and one may prey on the other?
- ... that populations of the endangered Albanian water frog (pictured) are declining due to over-collection for the food industry and pet trade?
- ... that the Sacramento Mountain Salamander is preyed on by the western terrestrial garter snake?
- ... that the female common rocket frog (pictured carrying tadpoles) and the male rainforest frog Colostethus stepheni are both involved in the parental care of their young?
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Caecilians (/sɪˈsɪliən/; New Latin for 'blind ones') are a group of limbless, vermiform (worm-shaped) or serpentine (snake-shaped) amphibians with small or sometimes nonexistent eyes. They mostly live hidden in soil or in streambeds, and this cryptic lifestyle renders caecilians among the least familiar amphibians. Modern caecilians live in the tropics of South and Central America, Africa, and southern Asia. Caecilians feed on small subterranean creatures such as earthworms. The body is cylindrical and often darkly coloured, and the skull is bullet-shaped and strongly built. Caecilian heads have several unique adaptations, including fused cranial and jaw bones, a two-part system of jaw muscles, and a chemosensory tentacle in front of the eye. The skin is slimy and bears ringlike markings or grooves and may contain scales.
Modern caecilians are a clade, the order Gymnophiona' /ˌdʒɪmnəˈfaɪənə/ (or Apoda /ˈæpədə/), one of the three living amphibian groups alongside Anura (frogs) and Urodela (salamanders). Gymnophiona is a crown group, encompassing all modern caecilians and all descendants of their last common ancestor. There are more than 220 living species of caecilian classified in 10 families. Gymnophionomorpha is a recently coined name for the corresponding total group which includes Gymnophiona as well as a few extinct stem-group caecilians (extinct amphibians whose closest living relatives are caecilians but are not descended from any caecilian). Some palaeontologists have used the name Gymnophiona for the total group and the old name Apoda for the crown group'. However, Apoda has other even older uses, including as the name of a genus of Butterfly making its use potentially confusing and best avoided. 'Gymnophiona' derives from the Greek words γυμνος / gymnos (Ancient Greek for 'naked') and οφις / ophis (Ancient Greek for 'snake'), as the caecilians were originally thought to be related to snakes and to lack scales.
The study of caecilian evolution is complicated by their poor fossil record and specialized anatomy. Genetic evidence and some anatomical details (such as pedicellate teeth) support the idea that frogs, salamanders, and caecilians (collectively known as lissamphibians) are each others' closest relatives. Frogs and salamanders show many similarities to dissorophoids, a group of extinct amphibians in the order Temnospondyli. Caecilians are more controversial; many studies extend dissorophoid ancestry to caecilians. Some studies have instead argued that caecilians descend from extinct lepospondyl or stereospondyl amphibians, contradicting evidence for lissamphibian monophyly (common ancestry). Rare fossils of early gymnophionans such as Eocaecilia and Funcusvermis have helped to test the various conflicting hypotheses for the relationships between caecilians and other living and extinct amphibians. (Full article...)List of selected amphibian type articles
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In the life cycle of a frog, a female lays her eggs in a shallow pond or creek, where they will be sheltered from the current and from predators. The eggs, known as frogspawn hatch into tadpoles. The tadpoles develop gradually into adolescent froglets and finally the froglet develops into an adult frog. -
Photo credit: Patrick CoinA terrestrial subadult Eastern newt or red eft (Notophthalmus viridescens). Salamanders of the family Salamandridae with aquatic adult stages are called newts. Some newts, including the Eastern newt, have a juvenile terrestrial stage called the eft. The red eft has bright aposematic coloration to warn predators of its highly toxic skin. -
Photo credit: Derek RamseyThe American Green Frog (Rana clamitans) is native to the eastern half of the United States and Canada. Green frogs live wherever there is shallow freshwater: ponds, road-side ditches, lakes, swamps, streams and brooks. The green frog is abundant wherever it occurs. -
Photograph: Benny TrappThe Albanian water frog (Pelophylax shqipericus) is a species of true frog in the family Ranidae. It is native to Albania and Montenegro, where it lives in aquatic environments. The frogs are medium-sized. Males sometimes bear a distinctive bright green stripe down the length of the backbone, but otherwise are green to light brown in overall colouring with large black or dark brown spots. Females are olive green or light brown in colour and also bear brown or black large spots. The species is endangered and known populations are currently in decline. -
Photo: QuartlDendrobates azureus, commonly known as the blue poison dart frog or okopipi, is a poison dart frog found in the forests surrounded by the Sipaliwini Savanna in southern Suriname. Authorities have recently treated it as a variant of Dendrobates tinctorius rather than a distinct species as before. -
Photograph: Benny TrappThe Epirus water frog (Pelophylax epeiroticus) is a species of frog in the family Ranidae. It is found in western Greece, including Corfu, and the southern areas of Albania. The species is collected from the wild for human consumption. -
The Blue Mountains Tree Frog (Litoria citropa) is a moderate-sized species of tree frog, up to about 60 mm (2.4 in) in length. It is native to coastal and highland areas of eastern Australia, especially in the Blue Mountains, hence its name. -
The edible frog (Pelophylax kl. esculentus) is a fertile hybrid of the pool and marsh frogs which is commonly found in Europe. The species is commonly used in food, including the French delicacy frog legs. -
Photograph: Benny TrappThe Iberian midwife toad (Alytes cisternasii) is a species of frog in the family Alytidae found in Portugal and western Spain. Its natural habitats are temperate forests and shrubland, Mediterranean-type shrubby vegetation, intermittent rivers and freshwater marshes, pastureland, and aquaculture ponds. The species is affected by habitat loss, and the IUCN lists it as being "Near Threatened". -
Photo credit: LiquidGhoulThe New England Tree Frog (Litoria subglandulosa) is a species of frog native to the streams of the New England Tablelands, New South Wales, Australia. -
Photo: Marie-Lan NguyenThe marsh frog (Pelophylax ridibundus) is a species of true frog and the largest frog native to Europe; females of this sexually dimorphic species may be up to 17 centimetres (6.7 in) long. The marsh frog feeds mainly on insects, but it also eats smaller amphibians, fish, and rodents. -
Photo credit: Fir0002The Eastern Banjo Frog (Limnodynastes dumerilli) is native to eastern Australia and has been introduced to New Zealand. The frog is also commonly called the pobblebonk after its distinctive "bonk" call, which is likened to a banjo string being plucked. Adults are roughly seven to eight cm long. -
A male Dendropsophus microcephalus frog, displaying its vocal sac—membranes of skin under the throat or on the corner of the mouth that distend to amplify its call. The species lives in moist areas in most of Central America and northern South America. -
The Eastern American Toad (Bufo americanus americanus) is a common subspecies of toad found throughout the eastern United States and Canada. It typically grows to 5–9 cm (2.0–3.5 in), with varying skin color and pattern depending on its environment. Its skin secretes bufotoxin, which is mildly poisonous to humans. -
Photograph: JJ HarrisonThe southern brown tree frog (Litoria ewingii) is a species of tree frog native to Australia found in a wide range of habitats. Reaching 45 millimetres (1.8 in), this species is generally brown, but green and green-striped morphs are also recorded.
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