Stygobromus

Stygobromus is a genus of amphipod crustaceans that live in subterranean habitats. The majority of the listed species are endemic to North America (one from Canada, the rest from the United States), a smaller number of species are also known from Eurasia. Most of the North American species live in areas which were not covered by the Laurentide Ice Sheet, although a few species seem to have survived under the ice. A number of species are on the IUCN Red List as endangered species (EN) or vulnerable species (VU); one species, S. lucifugus, is extinct.

Stygobromus includes the following species:


 * Stygobromus abditus Holsinger, 1978 (Virginia)
 * Stygobromus ackerlyi Holsinger, 1978 (Georgia)
 * Stygobromus alabamensis (Stout, 1911)
 * Stygobromus albapinus Taylor & Holsinger, 2011 (Nevada)
 * Stygobromus allegheniensis (Holsinger, 1967) (New York to West Virginia)
 * Stygobromus ambulans (F. Müller, 1846)
 * Stygobromus anastasiae Sidorov, Holsinger & Takhteev, 2010 (Siberia, Russia)
 * Stygobromus apscheronia (Derzhavin, 1945) (Azerbaijan)
 * Stygobromus araeus (Holsinger, 1969) Status iucn VU icon.svg (Virginia, North Carolina)
 * Stygobromus arizonensis Holsinger, 1974 Status iucn VU icon.svg (Arizona)
 * Stygobromus balconis (Hubricht, 1943) Status iucn VU icon.svg (Texas)
 * Stygobromus baroodyi Holsinger, 1978 (Virginia)
 * Stygobromus barri (Holsinger, 1967) Status iucn VU icon.svg (Missouri)
 * Stygobromus barryi Holsinger, 1978 (Tennessee)
 * Stygobromus behningi (Birstein, 1948)
 * Stygobromus bifurcatus (Holsinger, 1967) Status iucn VU icon.svg (Texas)
 * Stygobromus biggersi Holsinger, 1978 (Virginia to Pennsylvania)
 * Stygobromus blinni Wang & Holsinger, 2001
 * Stygobromus borealis Holsinger, 1978 (Vermont, New York)
 * Stygobromus boultoni Wang & Holsinger, 2001
 * Stygobromus bowmani (Holsinger, 1967) Status iucn VU icon.svg (Oklahoma)
 * Stygobromus canadensis Holsinger, 1980 (Alberta, Canada)
 * Stygobromus carolinensis Holsinger, 1978 (North Carolina)
 * Stygobromus chamberlaini Ellis, 1941
 * Stygobromus cherylae Wang & Holsinger, 2001
 * Stygobromus clantoni (Creaser, 1934) Status iucn VU icon.svg (Kansas, Missouri)
 * Stygobromus coeca (Dobraenu & Manolache, 1951)
 * Stygobromus coloradensis Ward, 1977
 * Stygobromus conradi (Holsinger, 1967) Status iucn VU icon.svg (Virginia)
 * Stygobromus cooperi (Holsinger, 1967) Status iucn VU icon.svg (West Virginia)
 * Stygobromus cowani Wang & Holsinger, 2001
 * Stygobromus culveri Holsinger, 1978 (West Virginia)
 * Stygobromus cumberlandus Holsinger, 1978 (Virginia)
 * Stygobromus curroae Wang & Holsinger, 2001
 * Stygobromus dejectus (Holsinger, 1967) Status iucn VU icon.svg (Texas)
 * Stygobromus dentata (Hubricht, 1943)
 * Stygobromus dershavini (Behning, 1928)
 * Stygobromus dicksoni Holsinger, 1978 (Alabama, Georgia)
 * Stygobromus donensis (Martynov, 1919)
 * Stygobromus duplus Wang & Holsinger, 2001
 * Stygobromus elatus (Holsinger, 1967) Status iucn VU icon.svg (Arkansas)
 * Stygobromus elliotti Holsinger, 1974
 * Stygobromus emarginatus (Hubricht, 1943) Status iucn VU icon.svg (Maryland, West Virginia)
 * Stygobromus ephemerus (Holsinger, 1969) Status iucn VU icon.svg (Virginia)
 * Stygobromus estesi Holsinger, 1978 (Virginia)
 * Stygobromus exilis Hubricht, 1943
 * Stygobromus fecundus Holsinger, 1978 (Tennessee)
 * Stygobromus fergusoni Holsinger, 1978 (Virginia)
 * Stygobromus finleyi Holsinger, 1978 (Tennessee)
 * Stygobromus flagellatus (J. E. Benedict, 1896) Status iucn VU icon.svg (Texas)
 * Stygobromus fontinalis Wang & Holsinger, 2001
 * Stygobromus franzi Holsinger, 1978 (Maryland)
 * Stygobromus gallawayae Wang & Holsinger, 2001
 * Stygobromus glacialis Wang & Holsinger, 2001
 * Stygobromus gracilipes (Holsinger, 1967) (Virginia to Pennsylvania)
 * Stygobromus gradyi Holsinger, 1974 Status iucn VU icon.svg (California)
 * Stygobromus grahami Holsinger, 1974
 * Stygobromus grandis Holsinger, 1978 (Georgia)
 * Stygobromus hadenoecus (Holsinger, 1966) Status iucn VU icon.svg (Texas)
 * Stygobromus harai Holsinger, 1974 Status iucn VU icon.svg (California)
 * Stygobromus hayi (Hubricht & Mackin, 1940) Status iucn EN icon.svg (District of Columbia)
 * Stygobromus herbsti Wang & Holsinger, 2001
 * Stygobromus heteropodus Hubricht, 1943 Status iucn VU icon.svg (Missouri)
 * Stygobromus hoffmani Holsinger, 1978 (Virginia)
 * Stygobromus holsingeri Ward, 1977
 * Stygobromus hubbardi Holsinger, 2009
 * Stygobromus hubbsi Shoemaker, 1942 Status iucn VU icon.svg (Oregon)
 * Stygobromus hyporheicus Wang & Holsinger, 2001
 * Stygobromus hyrcana (Dershavin, 1939)
 * Stygobromus idahoensis Wang & Holsinger, 2001
 * Stygobromus imperialis Wang & Holsinger, 2001
 * Stygobromus indentatus (Holsinger, 1967) Status iucn VU icon.svg (Maryland, North Carolina and Virginia)
 * Stygobromus inexpectatus Holsinger, 1978 (Alabama)
 * Stygobromus interitus Holsinger, 1978 (Virginia)
 * Stygobromus intermedia (Dobreanu, Manolache & Piascariu, 1952)
 * Stygobromus interstitialis Wang & Holsinger, 2001
 * Stygobromus iowae Hubricht, 1943
 * Stygobromus jakutana (Martynov, 1931)
 * Stygobromus jemezensis Wang & Holsinger, 2001
 * Stygobromus johanseni (Shoemaker, 1920)
 * Stygobromus kazakhstanica Kulkina, 1992
 * Stygobromus kenki Holsinger, 1978 (District of Columbia, Virginia)
 * Stygobromus lacicolus Holsinger, 1974
 * Stygobromus lanensis Wang & Holsinger, 2001
 * Stygobromus latus Wang & Holsinger, 2001
 * Stygobromus leensis Holsinger, 1978 (Virginia)
 * Stygobromus lepida (Mateus & Mateus, 1991)
 * Stygobromus levanidovae (G. Karaman, 1991)
 * Stygobromus limbus Wang & Holsinger, 2001
 * Stygobromus longidactylus (S. Karaman, 1929)
 * Stygobromus longipes (Holsinger, 1966) Status iucn VU icon.svg (Texas)
 * Stygobromus lucifugus (Hay, 1882) Status iucn EX icon.svg (Illinois)
 * Stygobromus mackenziei Holsinger, 1974 Status iucn VU icon.svg (California)
 * Stygobromus mackini Hubricht, 1943 (Tennessee to West Virginia)
 * Stygobromus meschtscherica (Borutsky, 1929)
 * Stygobromus mikhaili Sidorov, Holsinger & Takhteev, 2010 (Siberia, Russia)
 * Stygobromus minutus Holsinger, 1978 (Georgia)
 * Stygobromus montanensis Holsinger, 1974 (Montana)
 * Stygobromus montanus (Holsinger, 1967) Status iucn VU icon.svg (Arkansas)
 * Stygobromus morrisoni (Holsinger, 1967) Status iucn VU icon.svg (Virginia, West Virginia)
 * Stygobromus mundus (Holsinger, 1967) Status iucn VU icon.svg (Virginia)
 * Stygobromus myersae Wang & Holsinger, 2001
 * Stygobromus mysticus Holsinger, 1974
 * Stygobromus nanus Holsinger, 1978 (West Virginia)
 * Stygobromus nortoni (Holsinger, 1969) Status iucn VU icon.svg (Tennessee)
 * Stygobromus obrutus Holsinger, 1978 (Virginia)
 * Stygobromus obscurus Holsinger, 1974
 * Stygobromus onondagaensis (Hubricht & Mackin, 1940) Status iucn VU icon.svg (Arkansas, Kansas, Missouri and Oklahoma)
 * Stygobromus oregonensis Holsinger, 1974 (Oregon)
 * Stygobromus ozarkensis (Holsinger, 1967) Status iucn VU icon.svg (Arkansas, Missouri and Oklahoma)
 * Stygobromus parvus (Holsinger, 1969) Status iucn VU icon.svg (West Virginia)
 * Stygobromus pecki (Holsinger, 1967) Status iucn EN icon.svg (Texas)
 * Stygobromus pennaki Ward, 1977
 * Stygobromus philareti (Birstein, 1948)
 * Stygobromus phreaticus Holsinger, 1978 (Virginia)
 * Stygobromus pizzinii (Shoemaker, 1938) Status iucn VU icon.svg (District of Columbia, Maryland, Pennsylvania and Virginia)
 * Stygobromus pollostus Holsinger, 1978 (Virginia)
 * Stygobromus pseudospinosus Holsinger, 1978 (Virginia)
 * Stygobromus putealis (Holmes, 1909) Status iucn VU icon.svg (Wisconsin)
 * Stygobromus puteanus Holsinger, 1974
 * Stygobromus quatsinensis Holsinger & Shaw, 1987
 * Stygobromus rallus Wang & Holsinger, 2001
 * Stygobromus redactus Holsinger, 1978 (West Virginia)
 * Stygobromus reddelli (Holsinger, 1966) Status iucn VU icon.svg (Texas)
 * Stygobromus rudolphi Wang & Holsinger, 2001
 * Stygobromus russelli (Holsinger, 1967) (Texas)
 * Stygobromus saltuaris Wang & Holsinger, 2001
 * Stygobromus secundus Bousfield & Holsinger, 1981
 * Stygobromus sheldoni Holsinger, 1974
 * Stygobromus sierrensis Holsinger, 1974
 * Stygobromus simplex Wang & Holsinger, 2001
 * Stygobromus smithi Hubricht, 1943 Status iucn VU icon.svg (Alabama)
 * Stygobromus sparsus Holsinger, 1978 (Tennessee)
 * Stygobromus spinatus (Holsinger, 1967) Status iucn VU icon.svg (West Virginia)
 * Stygobromus spinosus (Hubricht & Mackin, 1940) (Virginia)
 * Stygobromus stegerorum Holsinger, 1978 (Virginia)
 * Stygobromus stellmacki (Holsinger, 1967) Status iucn VU icon.svg (Pennsylvania)
 * Stygobromus subtilis (Hubricht, 1943) Status iucn VU icon.svg (Illinois, Michigan and Missouri)
 * Stygobromus tahoensis Holsinger, 1974
 * Stygobromus tenuis (S. I. Smith, 1874) (Connecticut to Maryland)
 * Stygobromus trinus Wang & Holsinger, 2001
 * Stygobromus tritus Holsinger, 1974
 * Stygobromus urospinatus Wang & Holsinger, 2001
 * Stygobromus utahensis Wang & Holsinger, 2001 (Utah)
 * Stygobromus vitreus Cope, 1872
 * Stygobromus wahkeenensis Wang & Holsinger, 2001
 * Stygobromus wardi Wang & Holsinger, 2001
 * Stygobromus wengerorum Holsinger, 1974 Status iucn VU icon.svg (California)