Thynnidae

The Thynnidae (also known as thynnid wasps, flower wasps, or thynnid flower wasps) are a family of large, solitary wasps whose larvae are almost universally parasitoids of various beetle larvae, especially those in the superfamily Scarabaeoidea. Until recently, the constituents of this family were classified in the family Tiphiidae, but multiple studies have independently confirmed that thynnids are a separate lineage.

Description
Most species are small, but they can be up to 30 mm long. The females of some subfamilies (Diamminae, Methochinae, and most Thynninae) are wingless and hunt ground-dwelling (fossorial) beetle larvae, or (in one species) mole crickets. The prey is paralysed with the female's sting, and an egg is laid on it so the wasp larva has a ready supply of food. In species where both sexes are winged, males are similar in size to the females, but are much more slender. The males of species with wingless females, however, are often much larger than the females and have wings; the adults mate in the air, with the female carried by the male's genitalia. Adults feed on nectar and are minor pollinators. As some of the ground-dwelling scarab species attacked by thynnids are pests, some of these wasps are considered beneficial as biological control agents.

Taxonomy
The family has five extant subfamilies, which were previously placed in Tiphiidae before it was found to be non-monophyletic. Thynnidae genera are classified as follows:



Subfamily: Anthoboscinae Turner, 1912

 * Anthobosca Guérin-Ménéville, 1838
 * Anthosila Genise, 1985
 * Calchaquila Genise, 1985
 * Cosila Guérin-Méneville, 1838
 * Lalapa Pate, 1947
 * Odontothynnus Cameron, 1904
 * Paramyzine Berg, 1898

Subfamily: Diamminae Turner, 1907

 * Diamma Westwood, 1835

Subfamily: Methochinae Rohwer, 1916

 * Methocha Latreille, 1804
 * Pterombrus Smith, 1869

Tribe: Austromyzinini

 * Austromyzinum Brown, 1985
 * Cleftomyzinum Brown, 1985

Tribe: Myzinini

 * Myzinum Latreille, 1803

Tribe: Mesini Argaman, 1994

 * Mesa Saussure, 1892
 * Hylomesa Krombein, 1968

Tribe: Meriini Costa, 1858

 * Afromeria Boni Bartalucci, 2007
 * Allomeria Boni Bartalucci, 2007
 * Braunsomeria Turner, 1912
 * Iswara Westwood, 1851
 * Komarowia Radoszkowski, 1886
 * Lamprowara Boni Bartalucci, 2004
 * Macromeria Westwood, 1835
 * Meria Illiger, 1807
 * Meriodes Boni Bartalucci, 2007
 * Myzinella Guiglia, 1959
 * Notomeria Boni Bartalucci, 2011
 * Parameria Guérin-Méneville, 1837
 * Poecilotiphia Cameron, 1902
 * Tamerlanella Boni Bartalucci, 2004
 * Weerpaga Argaman, 1994
 * Zezelda Argaman, 1994

Tribe: Elaphropterini Kimsey, 1992

 * Ammodromus Guérin-Méneville, 1838
 * Argenthynnus Genise, 1991
 * Atopothynnus Kimsey, 1991
 * Brethynnus Genise,, 1991
 * Chrysothynnus Turner, 1910
 * Dolichothynnus Turner, 1910
 * Elaphroptera Guérin-Ménéville, 1838
 * Eucyrtothynnus Turner, 1910
 * Merithynnus Kimsey, 1991
 * Mesothynnus Kimsey, 1991
 * Spilothynnus Ashmead, 1903
 * Telephorornyia Guérin-Ménéville, 1838
 * Upa Kimsey, 1991
 * Zeena Kimsey, 1991

Tribe: Rhagigasterini Ashmead, 1903

 * Aelurus Klug, 1840
 * Curvothynnus Brown, 2010
 * Dimorphothynnus Turner, 1910
 * Eirone Westwood, 1844
 * Rhagigaster Guérin-Ménéville, 1838
 * Rugosothynnus Brown, 2015
 * Umbothynnus Brown, 2008

Tribe: Scotaenini Kimsey,1992

 * Anodontyra Westwood 1835
 * Glottynnus Genise, 1991
 * Glottynoides Kimsey, 1991
 * Kaysara Carnimeo & Noll, 2018
 * Ornepetes Guérin, 1838
 * Pampathynnus Carnimeo & Noll, 2018
 * Parelaphroptera Turner, 1910
 * Pseudelaphroptera Ashmead, 1903
 * Pseudoscotaena Carnimeo & Noll, 2018
 * Rostrynnus Genise, 1991
 * Scotaena Klug, 1810

Tribe: Thynnini

 * Acanthothynnus Turner, 1910
 * Aeolothynnus Ashmead, 1903
 * Agriomyia Guérin-Ménéville, 1838
 * Amblysoma Westwood, 1841
 * Ariphron Kimsey, 2007
 * Arthrothynnus Brown, 1996
 * Aspidothynnus Turner, 1910
 * Aulacothynnus Turner, 1910
 * Belothynnus Turner, 1910
 * Campylothynnus Turner, 1910
 * Catocheilus Guérin-Ménéville, 1842
 * Chilothynnus Brown, 1996
 * Deuterothynnus Brown, 2010
 * Doratithynnus Turner, 1910
 * Dythynnus Kimsey, 2001
 * Elidothynnus Turner, 1910
 * Encopothynnus Turner, 1915
 * Epactiothynnus Turner, 1910
 * Guerinius Ashmead, 1903
 * Gymnothynnus Turner, 1910
 * Hathynnus Kimsey, 2003
 * Heligmothynnus Brown, 2010
 * Hemithynnus Ashmead, 1903
 * Iswaroides Ashmead, 1899
 * Leiothynnus Turner, 1910
 * Lestricothynnus Turner, 1910
 * Lophocheilus Guérin-Ménéville, 1842
 * Macrothynnus Turner, 1908
 * Megalothynnus Turner, 1910
 * Neozeleboria Rohwer, 1910
 * Oncorhinothynnus Salter, 1954
 * Pentazeleboria Brown, 1983
 * Phymatothynnus Turner, 1908
 * Pogonothynnus Turner, 1910
 * Psammothynnus Ashmead, 1903
 * Rhytidothynnus Brown, 2008
 * Tachynoides Kimsey, 1996
 * Tachynomyia Guérin-Ménéville, 1842
 * Tachyphron Brown, 1995
 * Tmesothynnus Turner, 1910
 * Thynnoides Guérin-Ménéville, 1838
 * Thynnoturneria Rohwer, 1910
 * Thynnus Fabricius, 1775
 * Zaspilothynnus Ashmead, 1903
 * Zeleboria Saussure, 1867
 * Zythynnus Kimsey, 2001

Extinct genera
The following 4 genera are extinct members of Thynnidae:

Subfamily: Anthoboscinae Turner, 1912

 * †Architiphia Darling, 1990

Subfamily: Methochinae Rohwer, 1916

 * †Brachymethoca Zhang, 1989

Subfamily: Myzininae Borner, 1919

 * †Geotiphia Cockerell, 1906
 * †Lithotiphia Cockerell, 1906