Malus

Malus ( or ) is a genus of about 32–57 species of small deciduous trees or shrubs in the family Rosaceae, including the domesticated orchard apple, crab apples and wild apples.

The genus is native to the temperate zone of the Northern Hemisphere.

Description


Apple trees are typically 4 – talI at maturity, with a dense, twiggy crown. The leaves are 3 – long, alternate, simple, with a serrated margin. The flowers are borne in corymbs, and have five petals, which may be white, pink, or red, and are perfect, with usually red stamens that produce copious pollen, and a half-inferior ovary; flowering occurs in the spring after 50–80 growing degree days (varying greatly according to subspecies and cultivar).

Many apples require cross-pollination between individuals by insects (typically bees, which freely visit the flowers for both nectar and pollen); these are called self-sterile, so self-pollination is impossible, making pollinating insects essential.

A number of cultivars are self-pollinating, such as 'Granny Smith' and 'Golden Delicious', but are considerably fewer in number compared to their cross-pollination dependent counterparts.

Several Malus species, including domestic apples, hybridize freely.

The fruit is a globose pome, varying in size from 1 – in diameter in most of the wild species, to 6 cm in M. sylvestris sieversii, 8 cm in M. domestica, and even larger in certain cultivated orchard apples. The centre of the fruit contains five carpels arranged star-like, each containing one or two seeds.

Subdivisions and species
36 species and 4 hybrids are accepted. The genus Malus is subdivided into eight sections (six, with two added in 2006 and 2008). The oldest fossils of the genus date to the Eocene (Lutetian), which are leaves belonging to the species Malus collardii and Malus kingiensis from western North America (Idaho) and the Russian Far East (Kamchatka), respectively.

Species
36 species and four natural hybrids are accepted:


 * Malus angustifolia (Aiton) Michx. – southern crabapple
 * Malus asiatica Nakai – Chinese pearleaf crabapple
 * Malus baccata (L.) Borkh. – Siberian crabapple
 * Malus brevipes – shrub apple
 * Malus coronaria (L.) Mill. – sweet crabapple
 * Malus crescimannoi Raimondo
 * Malus daochengensis C.L.Li
 * Malus domestica (Suckow) Borkh. – domestic or orchard apple
 * Malus doumeri (Bois) A.Chev. – Taiwan crabapple
 * Malus florentina (Zuccagni) C.K.Schneid. – Florentine crabapple, hawthorn-leaf crabapple
 * Malus fusca (Raf.) C.K.Schneid. – Oregon or Pacific crabapple
 * Malus halliana Koehne – Hall crabapple
 * Malus honanensis Rehder
 * Malus hupehensis (Pamp.) Rehder – tea crabapple
 * Malus ioensis (Alph.Wood) Britton – prairie crabapple
 * Malus jinxianensis J.Q.Deng & J.Y.Hong
 * Malus kansuensis (Batalin) C.K.Schneid. – Calva crabapple
 * Malus komarovii (Sarg.) Rehder
 * Malus leiocalyca S.Z.Huang
 * Malus mandshurica (Maxim.) Kom. ex Skvortsov
 * Malus muliensis T.C.Ku
 * Malus niedzwetzkyana – Niedzwetzky's Apple
 * Malus ombrophila Hand.-Mazz.
 * Malus orientalis Uglitzk.
 * Malus prattii (Hemsl.) C.K.Schneid. – Pratt's crabapple
 * Malus prunifolia (Willd.) Borkh. – plum-leaf crabapple, Chinese crabapple
 * Malus rockii Rehder – native to China and Bhutan
 * Malus sikkimensis (Wenz.) Koehne – Sikkim crabapple
 * Malus spectabilis (Aiton) Borkh. – Asiatic apple, Chinese crabapple
 * Malus spontanea (Makino) Makino
 * Malus sylvestris (L.) Mill. – European crabapple
 * Malus toringo (Siebold) de Vriese (syns. Malus sargentii, Malus sieboldii) – Sargent crabapple, Toringo crabapple, or Siebold's crabapple
 * Malus toringoides (Rehder) Hughes – cut-leaf crabapple
 * Malus transitoria (Batalin) C.K.Schneid. – cut-leaf crabapple
 * Malus trilobata (Labill. ex Poir.) C.K.Schneid. – Lebanese wild apple, erect crabapple, or three-lobed apple tree
 * Malus turkmenorum Juz. & Popov (syn. Malus sieversii) – wild ancestor of cultivated species Malus domestica
 * Malus yunnanensis (Franch.) C.K.Schneid. – Yunnan crabapple
 * Malus zhaojiaoensis N.G.Jiang
 * Hybrids
 * Malus × floribunda Siebold ex Van Houtte – Japanese flowering crabapple
 * Malus × kaido (Wenz.) Pardé (syn. Malus × micromalus) – midget crabapple
 * Malus × soulardii (L.H.Bailey) Britton
 * Malus × zumi (Matsum.) Rehder

Formerly placed here

 * Macromeles tschonoskii (Maxim.) Koidz. (as Malus tschonoskii (Maxim.) C.K.Schneid.) – Chonosuki crabapple and pillar apple

Selected artificial hybrids

 * Malus × sublobata – yellow autumn crabapple (M. asiatica × M. toringo)

Fossil species
After
 * Malus collardii Axelrod, North America (Idaho), Eocene
 * Malus kingiensis Budants, Kamchatka Peninsula, Russia, Eocene
 * Malus florissantensis (Cockerell) MacGinitie Green River Formation, North America (Colorado) Eocene
 * Malus pseudocredneria (Cockerell) MacGinitie Green River Formation, North America (Colorado) Eocene
 * Malus idahoensis R.W.Br. North America (Idaho), Miocene
 * Malus parahupehensis J.Hsu and R.W.Chaney Shanwang, Shandong, China, Miocene
 * Malus antiqua Doweld Romania, Pliocene
 * Malus pseudoangustifolia E.W.Berry North America (South Carolina), Pleistocene

Cultivation
Crabapples are popular as compact ornamental trees, providing blossom in spring and colourful fruit in autumn. The fruits often persist throughout winter. Numerous hybrid cultivars have been selected.

Some crabapples are used as rootstocks for domestic apples to add beneficial characteristics. For example, the rootstocks of Malus baccata varieties are used to give additional cold hardiness to the combined plants for orchards in cold northern areas.

They are also used as pollinizers in apple orchards. Varieties of crabapple are selected to bloom contemporaneously with the apple variety in an orchard planting, and the crabs are planted every sixth or seventh tree, or limbs of a crab tree are grafted onto some of the apple trees. In emergencies, a bucket or drum bouquet of crabapple flowering branches is placed near the beehives as orchard pollenizers.

Because of the plentiful blossoms and small fruit, crabapples are popular for use in bonsai culture.

Cultivars
These cultivars have won the Royal Horticultural Society's Award of Garden Merit:-


 * 'Adirondack'
 * 'Butterball'
 * 'Comtesse de Paris'
 * 'Evereste'
 * 'Jelly King'='Mattfru'
 * 'Laura'
 * Malus × robusta 'Red Sentinel'
 * 'Sun Rival'

Other varieties are dealt with under their species names.

Toxicity
The seeds contain cyanide compounds.

Uses
Crabapple fruit is not an important crop in most areas, being extremely sour due to malic acid (which like the genus derives from the Latin name mālum), and in some species woody, so is rarely eaten raw. In some Southeast Asian cultures, they are valued as a sour condiment, sometimes eaten with salt and chilli or shrimp paste.

Some crabapple varieties are an exception to the reputation of being sour, and can be very sweet, such as the 'Chestnut' cultivar.

Crabapples are an excellent source of pectin, and their juice can be made into a ruby-coloured preserve with a full, spicy flavour. A small percentage of crabapples in cider makes a more interesting flavour. As Old English Wergulu, the crab apple is one of the nine plants invoked in the pagan Anglo-Saxon Nine Herbs Charm, recorded in the 10th century.

Applewood gives off a pleasant scent when burned, and smoke from an applewood fire gives an excellent flavour to smoked foods. It is easier to cut when green; dry applewood is exceedingly difficult to carve by hand. It is a good wood for cooking fires because it burns hot and slow, without producing much flame. Applewood is used to make handles of hand saws; in the early 1900s 2,000,000 board feet of applewood were used annually for this purpose.